The Nomenclature News of Flora Palaestina

Avinoam Danin

Department of Evolution, Systematics, and Ecology

The A. Silberman Institute for Life Sciences

The Hebrew University of Jerusalem

Jerusalem, Israel 91904

 

First published in 2000, in: Flora Mediterranea, 10: 109-172.

 

A note to the reader by A. Danin July 17, 2004

Dear reader,

When I wrote this article, at the end of 1999, I was ready to start working on the publication of my book listed below. As often happens, the preparation time has been longer than the wishful thinking of the author. I therefore opened the present path at the website of the Jerusalem Botanical Garden, in which I intended to update the readers in nomenclature changes. The big day of printing the book is close now. I am expecting to see the blue-prints of the book this week. I take the opportunity of the present stage in nomenclature changes for several years and make a renewed issue of the "Nomenclature News". The new additions are made in the same font type etc.  As a result of the changes the pagination became slightly different from the original publication in Flora Mediterranea.

Sincerely yours, A. Danin

A note to the reader by A. Danin January 9, 2005

Dear reader,

I was sorry to discover my mistake in naming Acacia salicina Lindl. As Acacia longifolia. Much of the information presented now under A. salicina was erroneously provided to A. longifolia. If the latter is found in Israel, it is not as common as Acacia salicina.  Sincerely yours, A. Danin

Abstract

During the preparation of the list of species in the Flora palaestina area several hundred names have been changed, added or replaced. Due to the limited space in the forthcoming book, the author lists the comments concerning many of the new names in Flora Palaestina.

Introduction

The material presented here is based on the material reported by the author in the manuscript and final proof-readings of “Distribution Atlas of plants in the Flora Palaestina area” which is part of the Flora Palaestina project, published by The Israel Academy for Science and Humanities. However, since the information in the “Atlas” is limited to the actual name of the plant and the name used in Flora Palaestina (Zohary, 1966, 1972; Feinbrun-Dothan 1978, 1986) there is no information as to why names were changed or added, where it is published etc. The aim of the present source of information is to provide, whenever available, tools for the reader to read for himself the information which led the author to the nomenclature conclusions that are presented in "the Atlas". Field observations on ecological conditions and history of invasion of established aliens are presented as well. The information presented on 19 species reported in Danin (2000) is incorporated in the present article as well.

Content

1.     List of the species “new to science” reported for the Flora Palaestina area.    p. 110

2.     List of species “new to” the Flora Palaestina area.                  p. 116

3.     List of species which were mis-identified in Flora Palaestina   p. 138

4.     List of species the names of which in Flora Palaestina are regarded at present as synonyms                                                                                            p.140

5. List of species, which were recorded in literature from the study area but are excluded from the “Distribution Atlas”                                                                               p. 151

6. References                                                                            p. 157

1. List of species “new to science” reported after the publication of Flora Palaestina

 

Allium daninianumBrullo, Pavone & Salmeri

Ref.: Brullo et al., (1996).

Fl.Pal.: “Allium stamineum“.

Note: The plants that were used to be called Allium stamineum Boiss. in Flora Palaestina (IV) and in earlier studies of that area were regarded by Brullo et al., (1993, in Candollea 48:279-290) as a complex. These authors regard the typical A. stamineum Boiss. to be confined to SW Turkey . Part of the complex named by Brullo et al., (1996) as the independent species depicting the cooperation in field trips of S. Brullo and A. Danin through OPTIMA expedition in Israel, 1989 (Bocconea 3: 5‑17).

Allium kollmannianumBrullo, Pavone & Salmeri

Ref.: Brullo et al., (1991).

Note: a rare plant, poorly collected, known from 3 locations or 3 specimens described by the authors. They regard it as confined to shrub-steppes on sandy-loess soils.

Allium tardiflorumKollmann, Shmida & Cohen

Ref.: Kollmann  et al., (1990).

Note: The only autumn-flowering Allium in Israel . Whereas all other species known from the area bloom in winter, spring, or early summer.

Amygdalus ramonensis Danin

Ref.: Danin, (1980).

Type specimen: Israel , Central Negev Highlands , Nahal Eliav, 4 km SW of Har Ramon, 34039’E/30038’N banks of wadi with loessial alluvium, 16.iv.1979, Danin (HUJ, E, K).

Note: a rare tree confined to rock crevices and other rocky mesophytic microhabitats in steppe-forests of the Negev Highlands (Danin, 1999a, 1999b).  A few specimens occur in banks of large wadis draining rocky terrain in the Negev Highlands. According to Browicz & Zohary (1996), who “summed up their views for tree breeders” (as D. Zohary explained the essence of their paper, pers. comm.) this taxon should be sunk into the complex of Amygdalus communis L. subsp. microcarpa (Post) Browicz & Zohary. I keep regarding this taxon as an independent species until more comprehensive study proves otherwise.

Hormuzakia negevensis (Danin) Danin & Hilger comb. nov.

Basionyme.: Anchusa negevensis Danin

Ref.: Danin (1995); Danin & Hilger (2000).

Type specimen:Israel , Negev Highlands , Yamin Plain, 10 km SE of Dimona, sandy soil, 21.3.1994 Danin (holo. HUJ; iso. E, K)

Note: This is the most rare endemic species known so far in Israel . It was not discovered, so far, outside its locus classicus, an area of ca. 1000 m2 of weathered sandstone, ca 10 km S of Dimona, suffering from constant erosion by westerly winds and rain-drops falling in the same direction as a result of the prevailing winds at rainfall events. It was discovered while the author looked for morphological diagnostic characters of desert psammophytes that grow in sites with constant wind erosion. Three other perennial desert Boraginaceae were known already, having the ability to produce shoots from wind-exposed roots (“Moltkiopsis ciliata type”, Danin, 1996). Such ability may protect the exposed roots from further exposure of the sand cover as the new adventitious stems locally decrease wind velocity and function as a sand trap. Hormuzakia negevensis shares the ability to produce root-borne shoots and grows on the rare combination of Neogene sandstone on the boundary of shrub-steppe and desert vegetation (Danin 1999a: p. 114; Fig. 5.13). Such a combination of old sandstone and climate may be searched for in S Jordan and Saudi Arabia , and H. negevensis may be found there.

          During the summer of 2000, as a result of DNA sequencing of herbarium specimen of this taxon by H.H. Hilger, as part of broader research on Boraginaceae, interesting findings were revealed. Our conclusions there were:” Analysis of the trnL(UAA)intron of chloroplast DNA corroborates the separate generic status of Hormuzakia Guşul. (Boraginaceae) apart from Anchusa L., and justifies the new combination of Hormuzakia negevensis (Danin) Danin & Hilger.

Artemisia jordanica Danin

Ref.: Danin (1999c)

Type specimen: S Jordan, Ma’an district, 45 km N of Mudawwara, sandy wadi, 26.10.1997, Danin, 972901 (HUJ; isotypes B, BM, E, K).

Note: This species was regarded by botanists dealing with the floras of Jordan and Saudi Arabia as Artemisia monosperma. However, the latter, having pinnatisect glabrous leaves (with rare hairy individuals), grows west of the Arava rift valley, the Gulf of Elat-Aqaba and the Red Sea whereas A. jordanica is the vicariant, with almost always entire leaves having appresed whitish indumentum, growing in mobile and stable sandy soils of S Jordan, Saudi Arabia, and Iraq.

Cyperus sharonensis Danin & Kukkonen

Ref.: Danin (1995).

Type specimen: Israel , Sharon Plain , 3 km S of Caesarea, 500 m N of Hadera Power Station, grid: 34053’15’’E/32028’20’’N, shallow dune on soil of ancient ruins, 27.5.1994, A. Danin (HUJ, H).

Note: This species was overlooked in the past being considered as part of the forms of Cyperus capitatus Vand., which grow sympatrically with C. sharonensis. Eig wrote in the 1930’s a note on one specimen calling attention to its peculiar morphology. The new species may be easily distinguished by the tall inflorescence (peduncles up to 100 cm long) and thick rhizomes (8-10 mm in diam.) of C. sharonensis contrary to the short peduncle (5-15 cm long) and thin rhizomes (1-2 mm in diam.).

Diplotaxis villosa Boulos & Jallad

Ref.: Boulos & Jallad (1975), Bot. Not.  128: 367.

Kickxia petrana Danin

Ref.: Danin (1991), Willdenowia 21: 181‑184.

Type specimen: Jordan, Edom, Aupstieg ed-Deir, 910 m, Ostern 1987, I. Künne 3488 (B, HUJ)

Note: After studying 3 other closely related species: K. acerbiana (Boiss.) Täckh. & Boulos, K. macilenta (Decne.) Danin, and describing K. judaica Danin, I suspected that the Kickxia species found in the sandstone terrain of Edom (SW Jordan ) differs from the other taxa in this group. Preparing scanning electron micrographs (SEM) of the seeds revealed its autonomous seed morphology.

Lathyrus golanensisCohen & Plitmann

Ref.: Cohen & Plitmann, in Plitmann et al., (1995): 250-252.

Note: The type population is from herbaceous vegetation on dark brown alluvial heavy soil, inundated and muddy in winter, very dry in summer.

Lathyrus plitmannii Greuter & Burdet

Ref.: Greuter & Burdet in Greuter & Raus (1989): p. 32.

Note: The authors discovered that the name L. lineatus given first to this taxon, by Plitmann & Heyn, is not valid and renamed it after one of its authors.

Lathyrus hirticarpusMattatia & Heyn

Ref.: Mattatia & Heyn (1976).     

Lens odemensisLadizinsky

Ref.: Ladizinsky (1986).                  

Note: Collected in Israel in herbaceous vegetation on basaltic slopes.

Micromeria danaensis Danin

Ref.: Danin  (1997).

Type specimen: Jordan , Edom , Dana Reserve, Wadi Barra area, 2 km SE of the visitor center. In crevices of smooth-faced white sandstone outcrops,  14.5.1996, A. Danin 963013 (holo HUJ; iso B).

Note: A narrow endemic growing in crevices of hard sandstone outcrops in steppes-forest area of SW Jordan (Danin 1999). It was not found so far out of Dana Nature Reserve. Its closest relative is Micromeria serbaliana Danin & Hedge, a narrow endemic of S Sinai , confined to crevices in smooth granite of a few mountain tops.

Origanum petraeum Danin

Ref.: Danin (1990).

Type specimen: Jordan , Edom , ca. 20 km S of Petra, Wadi Umm el Alla, 700-800 m, at the foot of sandstone cliff, 28.9.1988, I. Künne 21.8.4 (HUJ)

Note: Several additional populations of O. petraeum were discovered in the sandstone outcrops of SW Jordan , at elevation above 1000 m in Dana Nature Reserve, and north and south of Petra .

Origanum punonense Danin

Ref.: Danin, (1990).                        

Type specimen: Jordan, Karak (Edom ), Wadi Ghuweir, SE of Fenan (the Biblical Punon), 450 m, in sandstone crevices, 7.3.1986, Baierle & Kürschner 86-193 (BSB!).

Note: A few additional populations of O. punonense were discovered in the sandstone outcrops of SW Jordan , at elevation above 1000 m in Dana Nature Reserve, and north of Petra .

Origanum jordanicum Danin & Künne

Ref.: Danin & Künne, (1996).

Type specimen: Jordan: Edom, 8 km south of Petra,  1 km west of Et Taiyiba, Qseir el Anajil, 1180 m above sea level, coordinates: 30°26'E/30°15'N,, in crevices of Ordovician sandstone cliff, eastern exposure, 30.9.1994, I. Künne 94‑09‑19 (HUJ).

Note: A specimen collected at the NW slopes of Jebel Umm Adami, S. Jordan , (12.10.1998, Danin 981002) is more hairy than the type. Inquiry on vernacular names among Bedouin of Wadi Rum area reveal that they call this species ZA’ATARAN, whereas the name ZA’ATAR is applied to Satureja nabateorum Danin & Hedge. This means that there are more populations of O. jordanicum, unknown to scientists, which should be searched for and studied morphologically.

Paronychia jordanica Chaudhri      

Note: This species was published as a result of a study at the herbarium. The author (Chaudhri  1968) did not have a chance of observing populations of this species. He was not aware of the geography and named a plant growing in Nazareth , Israel (or if using an old geographical name “Palestine ”) after Jordan . This species should be thoroughly and critically investigated.

Polygala negevensis Danin

Ref.: Danin (1987)  .

Type specimen: Israel , southern Negev , 22 km north-northwest of Elat, crevices of hard limestone in a wadi, 1.7.1986, Danin (HUJ).

Note: Boulos (pers. comm. 8/2000) in his manuscript of the Flora of Egypt (in press) intends to sink this taxon into a synonym of the hairy species (with patent hairs) Polygala sinaica Botsch. When describing this taxon as an independent species I based my decision on plenty of morphological, ecological, and phytosociological observations in the field, which correlate perfectly with the morphology and ecology of the two taxa. I assume that Prof. L. Boulos could not have the opportunity to critically observe the plants in the field and based his decision on herbarium observations, which I am sure insufficient in this case. I keep the two taxa separate and independent at a specific rank.

Pycnocycla saxatilisDanin, Hedge & Lamond

Ref.: Danin et al., (2000).

Type specimen: Jordan, Edom: Rum area, 10 km SSE of Wadi Rum rest house, 35° 29' E 29° 31' N; in crevices of hard sandstone outcrops, N-facing, 1100 m, 12.10.1998, Danin 981101 (holo HUJ; iso E, B).

Note: Small populations of this species occur in crevices of the sandstone inslebergs


of Wadi Rum, S Jordan , and continue in the same habitat without any recognition of political borders in Saudi Arabia . Most of the year this plant looks like a stem assimilant and resembles Deverra triradiata in its general appearance.

Rubia danaensis Danin

Ref.: Danin (1997).

Type specimen: Edom , Dana Reserve, a limestone cliff above the visitor center, 13.5.1996, Danin 962509. (holo HUJ; iso B, E, K, W).

Note: Rubia danaensis is a narrow endemic (so far known) restricted to limestone cliffs of Edom - SW Jordan . It was found so far along a narrow strip less than 10 km wide and ca 60 km long at the edge of the S Jordanian plateau between Dana and Basta. It is closely related, morphologically, to the widely distributed vine of the Mediterranean maquis – Rubia tenuifolia D’Urv. Both species share the same slopes but differ in the habitat they grow, in functional morphology, and in phenology. Whereas R. tenuifolia may grow as a vine, climbing trees or shrubs on stony slopes or as a chasmophyte in crevices of limestone and sandstone, R. danaensis grows only as a chasmopyte on limestone cliffs. The differential morphological character is the teeth at leaf margins. Whereas the leaf margin of R. tenuifolia is always with pairs of sharp hooked teeth directed towards leaf base and towards its tip, the leaf margin of R. danaensis iscompletely smooth. The function of the hooked teeth is assistance to the climbing stems. If the two species look similar as derivatives of a common ancestor, the vine has a biological advantage by having hooked teeth. Smooth leaf margins may not harm the chasmophyte and having hooked teeth seems of no biological advantage in the limestone cliffs. Rubia tenuifolia blooms earlier than R. danaensis, and at first observations they seem to be isolated from each other phenologically by the flowering time.

Satureja nabateorum Danin & Hedge

Ref.: Danin & Hedge, (1998).

Type specimen: Jordan, Edom: 10 km E of Risha to Jebel Masú da, in crevices of smooth-faced white sandstone (Disi Formation), 650 m, 19.v.1997, Danin 970201 (holo HUJ; iso B, E, K)

Note: The closest relative of S. nabateorum is S. thymbrifolia Hedge & Feinbrun, described  in 1968 (both are in Sect. Zatarioideae Boiss.). The authors of S. thymbrifolia asked me, being the collector of the blooming holotype, to add a description of its habitats (Danin 1968). The discovery of the two species has a similar history; both were collected in the 1930’s as incomplete vegetative summer branches, which resembled each other. The resemblance of summer leaves of the two species caused the confusion and the inclusion of specimens of S. nabateorum in the description of S. thymbrifolia. The finding of fully developed plants of the second species led Danin & Hedge (1998) to recognize the significant morphological differences as sufficient between two species. The completely different ecological conditions, the long distance between the main populations of the two (150 km), and the morphological differences are further discussed by Danin & Hedge (1998). The occurrence of this species in adjacent location in Saudi Arabia , was cryptically reported in Danin et al., (2000). This report is based on specimens collected by Mrs. S. Collenette, observed by Mr. I.C. Hedge (pers. comm.), and deposited at the herbarium in Edinburgh (E). Their details are: Saudi Arabia: S of Tayma, Medinah road, shallow sand over white sandstone slabs, 3500 ft, Collenette 7814 (450 km SE of the southern collection site at Jebel Umm Adami in Jordan); off the Duba-Tabuk road, in crevices of white Rhum sandstone in red sand and thin basalt, Collenette 9059 (150 km SE of Jebel Umm Adami).

Scandix verna O.Cohen

Ref.: Cohen (2000).

Note: All the herbarium specimens of “Scandix iberica” from Israel were found to belong to the new taxon. However, one specimen from Edom seems to be the authentic S. iberica M.Bieb., and therefore the comment (p.p.) is presented in the Distribution Atlas.

Silene danaensis Danin

Ref.: Danin  (1997).

Type specimens: Jordan , Edom , Dana Reserve, a limestone cliff 500 m above and east of the visitor center 9.7.1996, Danin 963502 (holo HUJ; iso B, E, K).

Note: The moprphological similarity of S. danaensis to other species of Sect. Sclerocalycinae (Boiss.) Schischk. is compared and differentiated in the original description of the species (Danin 1997). S. danaensis grows in small extent in crevices of limestone outcrops. It is much more common in crevices of hard sandstone outcrops, where it grows from the area of Dana in the north, through most sandstone outcrops of Petra and Ras en Naqb area, to Jebel Umm Adami on the border of Jordan/Saudi Arabia and seems to grow in the Saudi Arabian sandstones although not reported from there yet.

Teucrium leucocladum Boiss. subsp. jordanicum Danin

Ref.: Danin, (1997).

Type specimens: Jordan : Edom , Dana Reserve, Wadi Barra, sandstone crevices, 9.7.1996, Danin 63601 (holo. HUJ, iso. B, E, K).

Note: The two most common species of Teucrium growing in semi-shrub communities of the bathas, steppes, and deserts of the Near East are T. capitatum L., which has forked or branched hairs, and T. leucocladum which has simple hairs. Both species have glandular hairs of various kinds. The typical subspecies of T. leucocladum is whitish and confined to warm lowlands of extreme desert areas in Israel , SW Jordan and S Sinai . subsp. jordanicum is brown-green and confined to rock crevices, mainly of sandstones in steppe areas. T. leucocladum subsp. sinaicum Danin (Danin  1997) is gray-green and grows in steppe areas of S Sinai , mainly in crevices of granite and other magmatic rocks. All the three subspecies have mostly globular inflorescences and hard branches, depending in the degree of their spinescence on the local grade of dryness.

Vicia basalticaPlitmann

Ref.: Plitmann (1987).

Note: Confined to moist soils on basalt.

 


2. Species “new to” the Flora Palaestina area

 

Acacia cyclops A.Cunn ex G. Don

Note:A plant of western Australian origin, planted in the arboretum near Kiryat Anavim as a possible afforestation tree. Following two events of “strong fire” in summers of 1987 and 1997, this presumably pyrophilous species became well-established on previously pine-afforested slopes. Following the two fire events this plant became one of the most prominent species in the area. It is assumed that additional wild fires in the Mediterranean mountainous area of Israel may open habitats for new establishment and possible naturalization of this species in Israel .

Acacia salicina Lindle. (common Australian names: willow acacia; willow wattle)

Note:This taxon was erroneously presented before as: Acacia longifolia (Andr.) Willd. Thanks are due to Prof. Alan Witztum for the correction of my mistake. A plant of eastern Australian origin, where it grows along water courses and flood plains. It is planted in settlements and urban gardens, mainly in the desert parts of Israel . There are a few habitats with poor competition where seedlings and trees, which established without human assistance are found. Most places are at the vicinity of seed source in ornamental areas of inhabited areas. In the extreme desert areas of the Arava Valley , establishment of A. salicina takes place in wadis, mainly near highways, where the ground was disturbed and water accumulation in the ground increase. Semi-spontaneous seedlings may populate irrigated areas with trickle-pipes in urban areas. Sandy soils in the sandy districts of the country in wadis near roads may support occasional trees as well.

Acacia paradoxa DC.

Note:A spiny shrub, introduced from Australia as an ornamental plant (Fahn, Heller & Avishai 1998). The collection details are: Israel , Judean Mts. , Kiryat Anavim, a rocky hill slope which suffered two wild fires in the last 20 years. 4.5.þ2004, Danin (HUJ, B, E).

When Kibbutz Kiryat Anavim was settled in 1920, one of the settlers’ activities was planting “The Arboretum” in cooperation with the Jewish National Fund. It was a rather large experimental plot and many Australian species were among the introduced species. From at least the 1950’s there was no additional planting. In the last 20 years there were two events of wild fire which led to a total elimination of the entire Arboretum and the pine forest planted on this slope. Today, more than 5 years after the last fire, there is a high proportion of Acacia cyclops A.Cunn ex G.Don,  A. saligna (Labill.) Wendl.f. in the arboreal components of the vegetation of the area which was “The Arboretum” long ago. Here and there are well-developed specimens of A. paradoxa. Finding it at roadside far away from the Arboretum site enable us to assume that it started to expand and should therefore be regarded as a “naturalized”. Authors: J.M. Dufour-Dror & A. Danin (submitted to Willdenowia June 2004)

Acacia saligna (Labill.) Wendl.f.

Ref.: Danin, in Greuter & Raus (1989): p. 29.

Note: A plant of western Australian origin, planted throughout the country as a fast growing shrub or tree. One of the main uses of the plant is in stabilization of the coastal sands. Pods with seeds derive from afforested areas and are blown by the wind. In the southern coastal plain, where dunes still exist, individual trees occur among sand dunes in habitats of sand deposition, mainly at the foot of the slip-face part of the dunes. This habitat is known also as “interdunes”. A. saligna trees are also planted in large quantities along roads in Israel .

Wild fires of afforested pine woodlands 30 km west of Jerusalem (Sha’ar HaGai area), where A. saligna trees were planted as well, took place in the late 1990’s. The nature of A. saligna as an aggressive colonizer (Stirton 1978) of burnt sites with low competition was prominent when the strong fire was followed by development of a green mantle of A. saligna seedlings in the first years after the fire. In summer 2000 this is the dominant plant of the slopes discussed above.

Spontaneously established A. saligna  trees are found also on crushed rocks which make up the sides of newly constructed roads, on fresh cuts of hills near roads, and on heaps of waste material derived from buildings.

Adonis flammea Jacq.

Ref.: Baierle (1993).

Note: The specimen collected by Baierle was determined by the expert for Adonis in the Near East , C.C. Heyn (BSB!)

Aegilops cylindrica Host

Ref.: Danin & Scholz, (1994).

Note: A small population of this adventive species was found in Jerusalem 500 m N of the entrance to the Givat Ram campus of the Hebrew University in the early 1990’s. A few months after the first finding most of the land on which it grew was removed in the process of construction of a new road. It was not collected or observed again yet.

Agrostis stolonifera L.

Note: The specimens in HUJ: Golan: Bab el Hawa, 25.7.1967, Y. Parag, Dan Valley : Tel Dan, near the Roman bridge, 24.6.1973. Both determined by Feinbrun.

Ailanthus altissima (Mill.) Swingle

Note: A tree introduced from China as an ornamental tree during the 20th century. Often planted in urban areas as ornamental, and easily escapes by vegetative propagation from roots and from seeds by germination and establishment in disturbed ground. It often germinates in crevices of sidewalks, walls of old buildings and roadsides, sends efficient root system below asphalt cover and breaks it. It is a dangerous invader that is hard to control.

Alisma gramineum Lej.

Ref.: Fragman, in Greuter & Raus (1999): p. 64.

Alliaria petiolata (M.Bieb.) Cavara & Grande

Note: The first specimens in HUJ: Judean Mts. , Jerusalem , The Hebrew University Campus, 15.3.1976 (possibly as an occasional alien); Golan, Tel Avital, 24.5.1987, Y. Or.

Aloe vera (L.) Burm.f.

Note: It is hard to state if the plant is an adventive synanthropic plant or spontaneous plant in the study area (cf. Shmida, 1995). It is known for a long time as a medicinal plant and could be brought here as such. The few populations found in Israel are confined to the proximity of Moslem cemeteries, where the A. vera was planted as ornamental. In Israel its flowers are mostly yellow.  In SW Jordan there is a large population of A. vera, with dark orange-coloured flowers, within the area of the ancient city of Petra . It is confined there to crevices in sandstone outcrops near the museum and looks rather spontaneous. A small population was found along the path of ancient road from Petra to Jebel Baida, some 3 km north of the main location within Petra . Further investigations in the sandstone desert of Jordan and of Saudi Arabia may prove the actual status of this species in the study area.

Alyssum desertorum Stapf

Ref.: Al-Eisawi (1985b).

Alyssum szowitsianum Fisch. & C.A.Mey.

Note: I could not find the origin of this record for Edom ; there are several herbarium specimens at HUJ from Mt. Hermon .

Amaranthus blitum L.

Note: This is a very common weed in Israel and Jordan at present. It might be that it was present in the study area for a long time, but was overlooked because of the high similarity to Amaranthus viridis L. (syn.: A. gracilis Desf.), which is known from the study area for a long time (Zohary 1966). The two diagnostic characters are, seed surface (completely smooth and shiny in A. blitum versus opaque and minutely punctate (in high magnification) in A. viridis. The fruit surface is slightly rugose and the fruit distinctly exceeds the perianth in A. blitum whereas strongly muricate, equaling or slightly longer than the perianth in A. viridis.

Amaranthus deflexus L.

Note: The first and only specimen in HUJ: Sharon , Hadera, orangerie, 7.10.1907, A.Aaronsohn. det. Liston.

Amaranthus rudis Sauer

Ref.: Danin & Liston, in Greuter & Raus (1986): 414. For additional aspects see Dafni & Heller (1990).

Note: This species is confined to naturally disturbed ground, albeit being part of a genus, most of species of which are known in the study area as colonizers of human-disturbed ground. It was collected mainly from banks of watercourses, artificial water reservoirs, or at the largest water reservoir of Israel – the Kinnereth (the Sea of Galilee ). In summer 2002 & 2004 it was observed growing as a weed in disturbed ground near roads in the Lower Galille (near Alonim and near Yifa’at).

Ambrosia confertiflora DC.

Ref.: Danin (1994).

Note: The population, which was sampled and discussed by Danin (1994), could not be detected after 1999. It is probably a case of non-successful establishment of an adventive plant.

Ambrosia tenuifolia Spreng.

Ref.: Danin (1994); Danin, in Greuter & Raus (1995): p. 172.

Note: The first specimen collected so far continue to multiply vegetatively in the first collection site: Israel , Sharon Plain : near Hadera, roadside in muddy soil, 27.12.1991, Danin (HUJ).

Ambrosia trifida L.

Note: The first specimen in HUJ: Akko Plain: Qishon bridge, 7.1987, Y. Marta.

Anabasis oropediorum Maire

Ref.: Danin, in Greuter & Raus (1982a): p. 38.

Andrzeiowskia cardamine Rchb.

Ref.: Heller & Liston in Greuter & Raus (1985): p. 63.

Note:  The first record in HUJ is: Golan: Dalhemiya, 7.4.1980, Y. Ivri.

Anoda cristata (L.) Schltdl.

Note: The first record in HUJ is: Pleshet, kibbutz Netzer Sereni, cotton field, July 1981, H.K. Mienis. For additional aspects see Mienis (1982) and Dafni & Heller (1990).

Arabidopsis thaliana (L.) Heynh.

Note: The first record in HUJ is: Golan: Masa’ada forest, 22.4.1973, Shmida, Dafni & Lerner. So far it was not collected elsewhere.

Araujia sericifera Brot.

Note: A vine, introduced from S Brazil as an ornamental plant in the late years of the 20th century. It escapes from cultivation and succeeds to establish itself in nearly abandoned citrus orchards and abandoned ones. It is wind-dispersed and being a vine it evidently has a biological advantage in germination and establishment at the shade of trees where not many competitors can survive. For additional aspects see Dafni & Heller (1990).

Arctotheca calendula(L.) Levyns

Specimen: Sharon, En HaKhoresh, a weed in irrigated young orange orchard on Red sandy-loam soil (Hamra), 29.03.2004, Danin (HUJ, B, E)

Note: This plant is a recent introduction from S. Africa to Israel . Agricultures in the Sharon told me that the plant was introduced by a soil-conservationist, working at the Israeli Ministery of Agriculture. His aim was to use the plant to protect soil erosion in sandy-loam soils, planted mainly by orange groves. It seems that a serious pest was introduced by this activity. It is a naturalized alien, expanding its distribution area every year. Authors: A. Danin & N. Kilian (submitted to Willdenowia June 2004).

Artemisia scoparia Waldst. & Kit.

Note: A few casual specimens of this un-naturalized alien are deposited in HUJ: Coastal Plain: Ramat Gan , 8.4.1936, Eig, Zohary & Feinbrun, N Negev : Beer Sheva, 31.10.1949, Y. D’Angelis.

Asparagus plumosus Baker

Note: A common cultivated vine in ornamental gardens. It succeeds very well as a vine in abandoned, almost abandoned, or slightly managed citrus orchards of the coastal plain. 

Asplenium scolopendrium L.

Ref.: Danin, in Greuter (1980a): p. 18.

Asplenium trichomanes L.

Note: According to Zohary (1976): p. 70, this species is known from the Galilee and the Golan. There are no herbarium specimens at HUJ to verify this note.

Astragalus brachystachys DC.

Ref.: Danin & Liston, in Greuter & Raus (1986): 427.

Astragalus camelorum Barbey

Ref.: Baierle (1993): p. 224. A. camelorum is a stem-assimilant semi-shrub rather easy to recognize by the medifixed hairs covering its stems and leaves. (cf. Danin 1976).

Astragalus eremophilus Boiss.

Ref.: Danin & Liston, in Greuter & Raus (1986): p. 427.

Astragalus ehrenbergii Bge.

Ref.: Baierle (1993): p. 224.

Atractylis mernephthae Asch., Schweinf. & Letourn. Note: First specimen in HUJ: Arava Valley : near Hazeva, sandy wadi, 1.1.1975, Danin, det. K. Schmid (M).

Atriplex holocarpa F.Muell.

Ref.: Danin, in Greuter & Raus (1982a): 40. Reported as Senniella spongiosa (F. Mueller) Aellen. For additional aspects see Dafni & Heller (1990).

Atriplex muelleri Benth.

Ref.: Danin, in Greuter & Raus (1982a): p. 38.

Atriplex nummularia Lindl.

Note:An indigenous Australian shrub, naturalized in California , introduced to Israel as a fodder during the 2nd half of the 20th century. There is but a few places where A. nummular established itself. However, since it resembles A. halimus in its general morphology it is possible that many individuals of A. nummularia were overlooked.

Atriplex suberecta Verd.

Ref.: Danin, in Greuter & Raus (1982b): p. 191.

Note: This species was first recorded by Danin (1982b) as A. microcarpa Benth. A more proper was found after the publication of the Chenopodiaceae in the Flora of Australia (Wilson 1984: p. 113).

Azolla filiculoides Lam.

Note: Specimens collected by M. Chaouat in the Golan without a specified location, nor with herbarium specimen were raised in the botanical garden at Mt. Scopus for a few years.

Barbarea brachycarpa Boiss.

Note: According to Zohary (1976): p. 156, known from the Golan.

Bassia scoparia (L.) A.J.Scott

Note: The first specimen at HUJ: Bet Shean Valley : Tirat Zvi, near ponds, 6.10.1970. For additional aspects see Dafni & Heller (1990).

Biarum auraniticum Mouterde

Ref.: Cohen (1999): p. 229.

 Note: The first specimen in HUJ: Golan, Rujm el Hiri, 2.11.1996, O. & M. Cohen.

Borrichia frutescens (L.) DC..,

Specimen: The first record in HUJ: Israel , the Philistean Plain, Tel Aviv coast, 1 km S of Reading Power Station. Crevices of side walk at the spray zone, 6.6.2000, Danin, det. D.J.N. Hind (K).

Note: A native of the USA , found from Virginia to Florida and Texas . It is apparently common on sandy sea-shores and in salt marshes. It has one obvious common name, of 'Sea Ox-eye'. The mode of its introduction into Israel is not known. It is not listed as an ornamental plant. It has a high potential of becoming an established alien. Being resistant to the sea-spray, evidenced by its origin and by the robust tufts with somewhat succulent leaves seen on the first collection site, there is a good chance that it will occupy more and more of this habitat in years to come.

Bothriochloa saccharoides (Sw.) Rydb.

Note: This plant is known so far only from the campus of the Hebrew University of Jerusalem, at Givat Ram, Jerusalem . It grows mainly as a weed in ornamental beds of small shrubs irrigated, but not frequently, in summer.

Brassica cretica Lam. subsp. aegea (Heldr. & Halácsy) Snogerup

Ref.: Heller & Liston in Greuter & Raus (1985): p. 64.

Note: The first record in HUJ: Carmel : Nahal Mearot, N exposure, cliffs, 10.6.1976, H. Lahav.

Brassica napus L.

Note: A common crop plant in Europe , where it is raised for its seeds, used for obtaining food oil. Thousands of tons of these seeds are imported by ship to Israel . Trucks transfer the seeds to the oil-factories destinations. Occasional seeds fall at the roadsides where no competitors exist as a result of roadside management practiced in Israel . It is impossible to call it an established alien, but it is constantly found at the roadsides.

Bromus chrysopogon Viv.

Ref.: Danin (1992): p. 72.

Bromus haussknechtii Boiss.

Ref.: Baierle et al., (1988):  p. 465

Bromus intermedius Guss.

Specimen seen: Lower Galilee , near Iblin, stony hills, 10.5.1967, Danin (HUJ), det. H. Scholz.

Bromus pseudobrachystachys H.Scholz

Ref.: Danin (1992): p. 74.

Bromus commutatusSchrad L. subsp. decipiens (Bomble & H.Scholz) H. Scholz

Note: In the first edition the opinion of Prof. H. Scholz was that the taxon we found in Israel deserves the name Bromus secalinusL. subsp. decipiens Bomble & H.Scholz. However, before the publication of the “Atlas” he wrote (pers. comm.) that he prefers now the new combination. In 1994 (Danin & Scholz 1994) he already placed the taxon in B. commutatus Schrad.

Bromus sericeus Drobov

Ref:  Danin (1992): p. 74.

Bupleurum libanoticum Boiss. & Blanche

Note: The first specimen in HUJ: Golan, between Buqata and Mas’ada, maquis on basalt rocks, 31.3.1967, M. Zohary, det. O. Cohen.

Calystegia soldanella (L.) Roemer. & Schult.

Note: First specimen in HUJ: Coast of Galilee , near Rosh Haniqra, 7.5.1976, Y.Cohen.

Campanula peregrina L.

Note: First specimen in HUJ: Upper Galilee, Nahal Keziv, Ein Tamir, 12.7.1985, A. Aaroni, det. O. Cohen.

Caralluma tuberculata N.E.Br.

Ref.: Baierle (1993): p. 215.

Castellia tuberculosa (Moris) Bor

Note: First record in HUJ: Dead Sea , low ground, alt. (-)394 m, 26.3.1912 Meyers & Dinsmore 72991; det. F. Scholz, 2000. Prof. H. Scholz discovered a small specimen of this rare species on a herbarium sheet of Bromus pseudobrachystachys H. Scholz.

Centaurea eriophora L.

Ref.: Witztum (1989).

Centranthus longiflorus Steven

Note: In moister Mediterranean countries, such as S France , Languedoc , and S Italy , this plant is a well-known colonizer of screes and other naturally disturbed habitats. In Israel it is planted in ornamental gardens, escapes from cultivation in urban areas and occasionally found in crevices in sidewalks and stone steppes in towns of the Mediterranean territories.

Chenopodium ficifolium Sm.

Ref.: Garve, in Greuter & Raus (1989): p. 28.

Chenopodium urbicum L.

Specimens examined: Israel : Kinnroth Valley , environs of Almagor, 8.9.1967, Danin & Zohary (HUJ!).

Ref.: Danin & Uotila, in Greuter & Raus (1989): p. 53.

Note: An important colonizer of the newly exposed shores of the coast of the Sea of Galilee , resulting from consequent drought years. Following several consequent dry years, this weed became the dominant and only species of many muddy areas, which were exposed in the long summer of 2000.

Chloris barbata Sw.

Specimen examined: Jordan, Ammon: Wadi Zarqa Ma’in, near the hot springs, an alluvial terrace, 18.xii.1997, Danin. 974301 (HUJ!), det. H. Scholz.

Note: The only place in the study area this plant is known of is the area near the hotel by the hot springs of Wadi Zarqa Ma’in. The plant grows in opened habitats of both natural and anthropogenic habitats. These are dry wadis that may flow temporarily in rainy years and in abandoned seasonal flowerbeds.

Chloris pycnothrix Trin.

Ref.: Danin & Cope, in Greuter & Raus (1995): p. 176.

Note: The only specimens available at the herbarium of the Hebrew University (HUJ) are those reported from the Philistean Plain from 1989.

Ciclospermum leptophyllum (Pers.) Sprague

Ref.: Danin, in Greuter & Raus (1995): p. 175.

Note:  Reported as Apium leptophyllum (Pers.) Benth., its older synonym from: Israel, Judean Mts. Jerusalem, irrigated lawn, 24.5.1990. This minute annual kept growing for additional two years in the small collection site and was not found there or anywhere else any more.

Cirsium vulgare (Savi) Ten.

Ref.: Liston, in Greuter & Raus (1987): p. 440.

Commicarpus sinuatus Meikle

Ref.: Greuter, in Greuter & Raus (1987): p. 448.

Note: Greuter (1987) writes that C. sinuatus is presented in Zohary (1966) from the southern Negev and Arava Valley as C. africanus var. viscosus.

Consolida coelesyriaca Mouterde

Note: Baierle (1993): p. 233.

Consolida hispanica (Costa) Greuter & Burdet

Note: The first specimen in HUJ: Golan, Qubet-Qarah, 6.5.1973, Didi & Edit. For nomenclature see Greuter in Greuter & Raus (1989): p. 43.

Consolida tomentosa (Aucher) Schrödgr. subsp. oligantha (Boiss.) Davis

Note: Boulos et al., (1975): p. 369.

Cornulaca monacantha Delile

Ref.: Danin, in Greuter & Raus (1982a): p. 39.

Corydalis triternata Zucc.

Note: The first specimen in HUJ: Golan, near Mas’ade, 1000m, maquis, 27.2.1968, M. Livneh.

Cotoneaster nummularius Fisch. & C.A.Mey.

Ref.: Baierle et al., (1988): p. 466.

Crassula vaillantii (Willd.) Roth

Ref.: Witztum & Raviv, (1988).

Cupressus semperinens L. var. sempervirens   

Note:  Mediterranean tree, domesticated from C. sempervirens L. var. horizontalis (Mill.) Aiton. Introduced as ornamental hundreds of years ago. The cultivar is found in the mountains of the Mediterranean territories of Israel near roads where extreme disturbance of the natural environment took place. Such places are parts of the roads where hills were cut in order to level mountains for preparing path for the asphalt road. Close source of seeds seems to be essential for semi-spontaneous establishment of the tree. There are a relatively higher number of trees in sites where rocks are of soft layers interbedded with fissured hard ones.

Another habitat where self-establishment of this variety of C. sempervirens takes place is abandoned quarries of limestone where heaps of crushed rocks occur. In the two habitats individuals of Cupressus sempervirens var. horizontalis and A. arizonica Green occur as well. The origin of their seeds is also from planted forests. However, in order to avoid confusion with the relict C. sempervirens var. horizontalis populations found in Edom , Jordan (Danin 1999) this taxon is not discussed here.

Dalbergia sissoo Roxb.

Ref.: Danin, in Greuter & Raus (1989): p. 31.

Note:The record of this plant from the waterfalls of En Gedi is based on flowers and leaflets I saw (and did not properly collect) floating on the water below the trees, which are extremely hard to access. After 1996 it was observed by Israeli visitors in a spring near Safi , Jordan , but not collected yet.

Damasonium polyspermum Coss.

Note: First specimen in HUJ: Lower Galilee , Kibbutz Khaquq, stagnant water, 22.4.1976, Rabinowitz.

Datisca cannabina L.

Ref.: Danin, in Greuter & Raus (1989): p. 28.

Note:No specimen of this species was collected in Israel after the report of the single specimen of this species reported as cited.

Digitaria ciliaris (Retz.) Koeler

Note: The first specimen in HUJ: Pleshet, Bar Ilan University campus, 20.9.1992, Y. Melamed.

Diplotaxis tenuifolia (L.) DC.

Ref.: Danin, in Greuter & Raus (1987): p. 440.

Note: In agreement with the note of 1987, this plant is not found yet out of Jerusalem area.

Ducrosia flabellifolia Boiss.

Note: Boulos & Al-Eisawi (1977b): p. 274.

Ehrharta erecta Lam.

Ref.: Danin & Scholz, in Greuter & Raus (1999): p. 65.

Specimens seen: Israel , Sharon Plain : Hanniel, 12 km E of Natanya, annual weed in irrigated orange grove, in the shade of trees, 10.4.1999, Danin.

Note: It is a recently introduced adventive weed, confined to sandy soils and partly shaded, summer-moist habitats.

Einadia nutans (R.Br.) A.J.Scott

Ref.: Danin & Heller, in Greuter & Raus (1998): p. 164.

Note: An indigenous lignified vine of S Australia and New South Wales , introduced as a potential ornamental plant for dry lands during the 2nd half of the 20th century. Escaped from experimental plots of plant introduction at a research institute in Beer Sheva, the N Negev . It has juicy and coloured small diaspores and is possibly distributed by small birds through endozoochory. It establishes itself in hedges mainly in the urban areas of Beer Sheva.

Enchylaena tomentosa R.Br.

Ref.: Danin & Heller, in Greuter & Raus (1998): p. 164.

Note: An indigenous subshrub to shrub of Australia , introduced as a potential fodder or an ornamental plant for dry lands during the 2nd half of the 20th century. Escaped from experimental plots of plant introduction at a research institute in Beer Sheva, the N Negev . Most individuals observed were in abandoned gardens in the urban areas of Beer Sheva. Its fruits resemble bird-dispersed diaspores, which may be dispersed through endozoochory.

Ephedra pachyclada Boiss. subsp. sinaica (H.Riedl) Freitag & Maier-Stolte (1994).

Note:The occurrence of E. pachyclada in the Near East was reported first from Sinai (Danin 1973: p. 25 and Danin & Hedge 1973: p. 264). I collected the type specimen of subspecies sinaica (H.Riedl) Freitag & Maier-Stolte in Sinai in smooth-faced granite cliffs. Riedl (1980) described Ephedra sinaica and based it on my specimens deposited in the herbarium at Edinburgh (E). Freitag & Maier-Stolte, (1994) sunk E. sinaica into a subspecies and reported a few specimens found in SW Jordan . It is rather common in steppes of Edom (SW Jordan ) on stony slopes.

Eragrostis virescens  J. Presl

Syn.:  Eragrostis mexicana (Hornem.) Link subsp. virescens  (J. Presl) Koch & Sanchez.

Note: The first specimen was collected in Jordan , at the lowlands of Edom : Ain Fidan, 40 km S of Safi, near a stream of fresh water, 17.7. 2000, Danin 2K0316, det. H. Scholz  (HUJ, B). It is a S American species in origin, but now distributed to warmer countries throughout the globe (Martin & Scholz, in Willdenowia 28:59-63 [1998]).

Eucalyptus camaldulensisDehn.    

Note: A tree introduced from W. Australia as an ornamental- and forest-tree at the 1880’s. Rarely germinating and establishing itself spontaneously in Israel. For many years E. camadulensis was one of the most common trees used in afforestation on various soils types. At its area of origin it dominates riparian vegetation (its common name in Australia is “river red gum”). The common habitat where germination and establishment of this tree takes place is ditches along the highway in areas of deep clayey soil (Grumusol) where irrigated agriculture is practiced near the road. Functioning as drainage canals, the ditches may have wet ground throughout the summer. Meeting the germination demands of E. camaldulensis in this habitat may be a result of the existence of moist soil in summer. Millions of seeds produced by the adult trees every year are wind-disperses.     

A nearly natural establishment of E. camadulensis took place in watercourses of the N. Negev following the rainy year 1990-91. In that year much of Israel , including the northern Negev received ca 200% the average annual rainfall. During the following three years wadis, which were sometimes streaming in winter drained water the year round. In such wadis E. camadulensis germinated and established themselves spontaneously as well.

Euphorbia bivonae Steud. Var. sianica Hadidi

Ref.: Al-Eisawi et al., (1996) recorded it from rocky ground, Wadi Rum, S Jordan.

Euphorbia cyathophora Murr.

Specimen: Dead Sea area: En Gedi, in a date palm plantation, irrigated with trickle-pipe system. 8.4.2004, Danin (HUJ, B, E) Det.: Dr. Mark Mayfield (Kansas State University ).

Note: The potential of En Gedi to become a site for the establishment of alien plants is rather high. It is one of the warmest sites in Israel and most of its agriculture is based on irrigation. This combination of high temperatures and water availability is rare in the rest of the country. Euphorbia cyathophora, which could have been brought to Israel as an ornamental plant, is subtended by large stands of the invasive naturalized alien Lantana camara L. The agricultures of Kibbutz En Gedi and the rangers of the Nature Reserve put much efforts to eradicate both invaders. (Submitted to Willdenowia June 2004).

Euphorbia lasiocarpa Klotzsch, E. serpens Kunth, and E. supina Rafin.

Note: The three species of Euphorbia listed above belong to the group of adventive and naturalized species which develop mainly in summer, in garden plots irrigated with trickle pipes. Thanks are due to Mr. Radcliff-Smith, at Kew , for the determination of these species in the 1990’s. That information was used by Feinbrun-Dothan & Danin (1991). For additional aspects see Dafni & Heller (1990).

Fagonia schimperi Presl

Specimen examined: Jordan , Edom : Rum area, 8 km  SE of Wadi Rum rest house, in crevices of smooth-faced granite outcrops, facing west, 13.10.1998, Danin 981304.

Ref.: Danin, in Greuter & Raus (1999): p. 64.

Note: This is one of the Fagonia  species already known from Sinai and discovered recently from Jordan . In both areas it is confined to rocky terrain, magmatic in Sinai and sandstone in Jordan .

Fallopia convolvulus (L.) Á.Löve

Ref.: Danin, in Greuter & Raus (1989): p. 41.

Note: An adventive weed, which grows in small quantities here and there in summer-irrigated ground.

Galium ghilanicum Stapf.

Ref.: Danin (1992): p.110.

Galium spurium L.

Ref.: Danin & Liston, in Greuter & Raus (1984b): p. 308.

Gamochaeta pensylvanica (Willd.) Cabrera

Ref.: Kilian & Danin, in Greuter & Raus (1999): pp. 53-54.

Specimens examined: Israel , Sharon Plain : Hanniel, 12 km E of Netanya, orange orchard,  21.4.1985, Raviv (HUJ!); Philistean Plain: Beerot Yitzhak, 24.4.1988, Flint (HUJ!).

Note: The ecological niche this adventive plant is occupying at present is disturbed, irrigated and shaded sandy soils of the coastal plain.

Gastridium phleoides (Nees & Meyen) C.E.Hubb.

Ref.: Scholz, in Greuter & Raus (1998): p. 171.

Note: In addition to the specimen reported by Scholz from the Negev Highlands I collected, March 2000, additional specimens of this species (det. H. Scholz) at the Judean Mts., 15 km W of Jerusalem (HUJ, B).

Geranium libanoticum Schenk

Note: The first specimen in HUJ: Golan, east of Buq’ata, NE face of Har Varda, volcanic tuff, alt. 1160 m, 5.5.1991, O. Cohen.

Hedysarum cf. pogonocarpum Boiss.

Ref.: Baierle et al., (1988): p. 463.

Note: The plant recorded and illustrated by Baierle et al., (1988) is evidently new to the Flora Palaestina area. Its final determination anticipates further investigation.

Helianthus annuus L.

Note: The first specimen in HUJ: Bet Shean Valley , near Tirat Zevi, roadside near fishponds, 8.7.1971, M. Zohary & A. Shmida. The identity of the adventive Helianthus species in the Flora Palaestina area is not critically reviewed yet. For additional aspects see Dafni & Heller (1990).

Heliotropium lasiocarpum Fisch. & C.A.Mey.

First specimen: Jordan, Ammon: 25 km S of Amman, Loessial soil near the road 1 km N of Queen Alia Airport, 16.7. 2000, Danin 2K0102; det.: H. Foerther (M) and Nadja Diane (BSB).

Henrardia pubescens (Bertol.) C.E. Hubbard

Ref.: Baierle et al., (1988): p. 465

Hippocrepis biflora Spreng.

Note: Prof. P. Lassen  (pers. comm., 2000) enlightened the following specimens determined by him: Sharon, Meyers & Dinsmore 6446 (E p.p.!); Upper Galilee : W. el Qarn, Pabot s. n. (G!); Esderaelon Plain, 23.4.1931, Naftolsky (HUJ!); Davis 4153 (E!, K!); Moav, Meyers & Dinsmore M446 (G!, K p.p.!).

Hohenackeria exscapa (Stev.) Kos.-Pol.

Ref.: Al-Eisawi et al., (1994).

Hypericum olivieri (Spach) Boiss.

Note: Boulos et al., (1975): p. 368.

Hypericum perforatum L.

Ref.: Liston, in Greuter & Raus (1987): p. 441.

Hypericum sinaicum Hochst. & Steud. ex Boiss.

Specimens seen: Edom , Dana Reserve, a limestone cliff 500 m above and east of the visitor center 13.5.1996, Danin 962609.

Note:Danin (1997) Willdenowia 27: p. 172. The population discovered in Jordan is very small. However, the limestone cliffs of SW Jordan are poorly investigated and it may be more common. The large distance of hundreds of kilometers between the populations known so far in Saudi Arabia and in Sinai may be regarded as discontinuity related to dispersal in the past when conditions enabled more continuous growth. Many species growing with H. sinaicum in Jordan are relicts of this kind (Danin, 1999a, 1999b).

Ipomoea hederacea Jacq.

Note: The first specimen in HUJ: Dan Valley , Kibbutz Dafna, in cotton field, 20.91984, D. Yoel & A. Liston.

Ipomoea indica (Burm.f.) Merr.

Ref.: Liston, in Greuter & Raus (1987): p. 440.

Note:  Liston (1987) mentions a specimen collected by M. Zohary 19.3.1962 and states that the status of that species is not known. The status did not change to the present day.

Ipomoea pes-caprae (L.) R.Br.

Note: Zohary (1976: p. 300) mentions the finding of this species in the coastal plain without an accurate location. There is no specimen of this species from the Flora Palaestina area in HUJ.

Ipomoea triloba L.

Note: Dan Valley , Kibbutz Dafna, in cotton field, 20.91984, D. Yoel & A. Liston.

Lactuca aculeata Boiss.

Note: First specimen in HUJ: Golan, near Nov, 23.9.1977, Golberson.

Lantana camara L.

Note: An ornamental plant introduced from Tropical America at an unknown date but, following the general history recorded by Stirton (1978), it seems to have been introduced during the 20th century. L. camara is a noxious weedy plant, which is rated as one of the ten worst weeds in the world (Stirton 1978). It is dispersed by birds through endozoochory and easily establishes itself in cultivated crops that are not intensively managed. A prominent habitat of this plant is almost abandoned or abandoned citrus groves. L. camara is often transferred from ornamental gardens to these abandoned orchards by birds. The latter tend to stand on trees and drop seeds, which passed in their intestine on soil poor in competing annuals, due to low light intensity. The weedy L. camara germinates and establishes itself in this habitat. L. camara often looks like a vine covering entirely dead citrus trees. Dafni & Heller (1990) discuss aditional aspects of this species.

Another common habitat of L. camara is datepalm plantations irrigated by trickle-pipes. These are common along the Jordan-Dead Sea-Arava rift valley. In this area and especially in the oasis of En Gedi it threats to outcompete local flora in natural habitats. In addition to these two clear habitats, L. camara is a common component of the wasteland vegetation in the lowlands of the Mediterranean territories of Israel .

Lappula barbata (M.Bieb.) Gürke

Specimen examined:Edom , northern section of Dana Reserve, the Cupressus sempervirens reserve, sandy soil 15.v.1996, Danin 963106. Reported by Al-Eisawi (1983) from the same location.

Lappula sinaica (DC.) Asch. & Schweinf.

Ref.: Baierle et al., (1988):  p. 460.

Note: A common steppe and desert annual of S Sinai . Collected without a clear record in Elat area and in a steppe-forest of Juniperus phoenicea in Edom north of Wadi Musa.

Lasiospermum brachyglossum DC. var. sinaicum Asch. & P. Hoffm.

Ref.: Al-Eisawi et al., (1996).

Lathyrus annuus L.

Note: The first specimen in HUJ: Sharon, E of Ma’agan Mikhael junction, abandoned field, 12.4.1994, Leon 71403.

Lathyrus ciliolatus Rech.f.

Ref.: Mattatia (1977).

Lathyrus palustris L.

Note: A single population was recorded from the Jerusalem Forest near Upper Motza . Possibly escaped from cultivation and established itself at the shade of forest trees and some wastewater derived from houses above it.

Lathyrus sphaericus Retz.

Ref.: Mattatia (1974).

Lavandula dentata L.

Ref.: Baierle et al., (1988): p. 463.

Lavatera arborea L.

Note: Occasional individuals develop on highly disturbed ground, mainly in urban areas and near roads.

Leopoldia  tenuiflora (Tausch) Heldr.

Ref.: Baierle et al., (1988): p. 463.

Leptochloa mucronata (Michx.) Kunth

Note: Danin & Scholz, in Greuter & Raus (1998): pp. 171-172, reported the occurrence of this species under its synonym:Leptochloa filiformis(Lam.) P. Beauv. For information concerning this synonymy see: Taxon 42: 413-417 (1993).

Leptochloa uninervia (J.Presl) Hitchc. & Chase

Ref.: Danin & Scholz, in Greuter & Raus (1999):  p. 66.

Note: This new adventive grass is confined to muddy soil of the newly exposed land at the beach around the Sea of Galilee . In the last few years, as a result of frequent droughts there is a considerable drop of water level of the lake during the summer. L. uninervia became one of the common colonizers of the expending land of moist soils of the beach.

Lotus hispidus DC. in Lam., Fl. Fr. ed. 3, 5 (=suppl.): 572 (1815)

Ref.: Lassen, in Greuter & Raus (1987): p. 445 (as L. subbiflorus Lag.; Greuter et al., (1989): p. 133.

Note: Lassen (1987) saw and determined two herbarium specimens from Israel , the Sharon Plain (pers. comm.); Greuter et al., (1989) presented the epithet L. hispidus DC. (1805) suggesting that this is the correct specific epithet. However, Lassen does not agree with this statement (in litt., Sept. 2000) and communicated that whereas L. hispidus DC. (1805) is a mistake, De Candolle corrected the mistake in 1815 edition.

Maireana brevifolia (R.Br.) P.G.Wilson

Ref.: Danin, in Greuter & Raus (1982a): p. 39.

Note: An Australian semishrub to shrub, introduced as an ornamental or fodder plant for dry lands during the 2nd half of the 20th century. Escaped from cultivation and established in roadsides where plants enjoy extra water supply as runoff from asphalt roads and low competition. It is also found in highly disturbed sites where soil was removed in the processes of road construction. The sites where it grows are mainly on slightly saline loessial soils.

Medicago arborea L.

Ref.: Baierle et al., (1988): p. 461.

Melia azedarach L.

Ref.: Danin, in Greuter & Raus (1989): p. 36.

Note: An ornamental tree of a SE Asian origin (the common names China-Tree or China-Berry may indicate its origin) which succeeds to establish itself in human disturbed habitats throughout the moist parts of the country. This tree produces fruits successfully wherever it grows. The fruits are bat-dispersed and the ground below fruit bats “resting” places may be covered by huge quantities of M. azedarach stones, each containing many small seeds. M. azedarach trees may be found in mal-managed and abandoned citrus groves. Wasteland, where competition is low for a long period of time, also supports this tree.

Melica canescens (Regel) Lavr.

Ref.: Baierle et al., (1988): p. 465

Mentha suaveolens Ehrh.

Note: Specimens collected by M. Chaouat in the Golan without a specified location nor with herbarium specimen are raised in the botanical gardens of Jerusalem at Givat Ram and Mt. Scopus .

Minuartia intermedia (Boiss.) Hand.-Mazz.

Ref.: Baierle et al., (1988): p. 460.

Minuartia sinaica (Boiss.) Danin

Ref.: Danin (1987).

Note: This taxon was regarded as a synonym of Minuartia picta (Sm.) Bornm. (Greuter et al., 1984: p. 223). When the two taxa meet together (e.g., the Negev Highlands in a good rainy year – cf. Danin, 1987), they grow on different soil types and differ morphologically. Their petal size and their leaf and stem indumentum correlates with seed surface morphology (using SEM images). These are not geographical races, nor local variation in certain populations, but a constant assemblage of morphological and ecological conditions, and therefore deserve a species level recognition.

Moenchia erecta (L.) P.Gaertn. et al.

Note: The first specimen in HUJ: Golan, clearing in forest near Mas’ada, basalt soil, 29.4.1968, M. Zohary.

Morus albaL.

Note: A fruit tree, introduced from C & E China. It was planted in some places as food for butterfly caterpillars (“silkworms”). Birds through endozoochory frequently disperse Morus alba. A preferred position for birds is high places; thus standing and watching the area around them, birds often use orchard trees as their watching sites and enrich the soil below them with droppings. Whereas the area below the fruit trees is ploughed against weeds, the abandoned orchards, during the first years after cultivation ceased are not ploughed and the seeds transported and dropped by the birds germinate and the plants establish themselves.

Myosotis refracta Boiss.

Note: The first specimen in HUJ: Jordan, 4 km S of Wadi Sabat, 2 km N of Jebel Umm Adami, 55 km ESE of Aqaba, 10.v.1998, Danin 980313.

Myosurus minimus L.

Note: The first specimen in HUJ: Upper Galilee , Har Meron, Khirbet Zabad, 1.5.1966, Sagi.

Najas guadalupensis (Spreng.) Magnus

Ref.: Witztum & Chaouat (1991): p.65-66.

Nicandra physalodes (L.) Gaertn.

Note: The first specimens in HUJ: Akko Plain, Haifa Bay , entrance to Shemen Factory, 17.6.1986, Y. Marta. Dafni & Heller (1990) discuss additional aspects.

Garidella nigellastrum L.

Note: Danin, in Greuter & Raus (1989): p. 44. See also the comment on Garidella unguicularis Poir.

Nigella segetalis M.Bieb.

Note: Collected once in the Negev Highlands. In that population, which developed on loessial soil of an ancient agricultural terrace between Nahal Elot and Borot Lotz, in the shrub-steppe zone at an elevation of approximately 1000 m, there were young specimens. Mature specimens should be collected there to make this record clear beyond any reasonable doubt.

Nitraria schoberi L.

Note: Recorded by Kislev et al., (1992) from 19,000 years old site at the shores of the Kinnereth (Sea of Galilee ). At present does not grow in the Flora Palaestina area, but east of the 360 longitude.

Nothoscordium inodorum (Aiton) Nicholson

Specimen: Israel , Judean Mts. : Bet HaKerem, a weed in irrigated flower garden. 17.05.2004,  Danin (HUJ).

Note: This xenphyte of American origin, seen in Israel for several years in irrigated ornamental plots in several locations, is first reported here. It was determined and compared to well-studied material in Cyprus by G. Hadjikyrikou. (Submitted to Willdenowia June 2004).

Oenothera biennis L.

Note: The first and only specimen in HUJ: Pleshet, 0.5 km S of Masmiya junction, 25.8.1989, Dafni.

Oenothera laciniata Hill

Note: Liston, in Greuter & Raus (1987): p. 448.

Oenothera rosea Aiton

Ref.: Danin & Shmida, in Greuter 1980: p. 232.

Ophioglossum polyphyllum A.Braun

Note: The first specimen in HUJ: Arava Valley , Hatzeva, Nahal Shezaf, sands, 11.3.1981, Shmida & Dagan.

Ophrys flavomarginata (Renz) H.Baumann & Künkele

Note: Dafni et al., (1987): p. 146.

Opopanax hispidus (Friv.) Griseb.

Note: Al-Eisawi (1983): p. 360.

Panicum coloratum L.

Ref.: Danin & Scholz in Greuter & Raus (1998): p. 172.

Papaver decaisnei Hochst. & Steud. ex Elkan

Note: Recorded from Jordan without specific location by Miller & Cope (1996): p. 344.

Papaver glaucum Boiss. & Hausskn.

Note: Boulos et al., (1975): p. 368.

Papaver somniferum L.

Note: There are occasional individuals of the cultivar growing at the roadsides of the mesic districts of the country. Their origin seems to be occasional seeds which fall-off of the condiment “papaver seeds” transported by vehicles. The individuals are found in the belt of 3-5 m along the road sprayed with herbicides and support many other cultivars, e.g. Secale cereale, Hordeum vulgare, Triticum aestivum and many other plants mentioned in the present paper.

Parapholis marginata Runem.

Note: First record: Israel , the Philistean Plain, Tel Aviv coast, 1 km S of Reading Power Station. Steep sandy slope at the spray zone, 6.6.2000, Danin, det. H. Scholz

Parkinsonia aculeata L.

Ref:Danin, in Greuter & Raus (1989): p. 34.

Note: A central-American tree, introduced during the 20th century as afforestation tree for dry lands. It is planted here and there as an ornamental tree all over Israel . It became one of the most aggressive colonizers at sides of roads sprayed with herbicides. It is growing as a dominant tree in savannas in Nicaragua and Costa Rica in deep clayey soils (known as Grumusols or Vertisols; Danin 1978). In Israel it constitutes rather dense stands on clayey soils, e.g., near the highway from Yagur to Tivon and east of Ben Gurion Airport , on the way to Jerusalem . Additional common habitats are wastelands, sites of demolished buildings material, and garbage collection heaps. In all these sites, and at the roadsides, poor competition to the establishing seedling prevails for some time.

Paronychia capitata (L.) Lam.

Note: The specimen determined by the monographer (Chaudhari 1968) is: Judean Mts. , Jerusalem , Matzleva, 6.5.1953, S. Leinkram. An older specimen is from Judean Mts., environs of Hartuv, 7.3.1924, Eig, det. Danin 1968. Zohary (1966): p. 131 thinks those earlier records of P. capitata from Flora Palaestina area “are most probably erroneous”. However, most of Zohary’s records of P. sinaica Fresen. from the Mediterranean territories of the study area seem to be of P. capitata.

Paronychia macrosepala Boiss.

Note: Coast of Galilee , S of Gesher Haziv, sand hill, 3.5.1956, Orshan, det. M.N. Chaudhari 1968. This is the only specimen of this species in HUJ.

Parthenium hysterophorus L.

Note: The first specimen in HUJ is: Bet Shean Valley , Tirat Zvi, 28.8.1979, Shmida. For additional aspects see Dafni & Heller (1990).

Passiflora morifolia Masters

Note: Joel &  Liston (1986): p. 219.

Peltaria angustifolia DC.

Ref.: Baierle et al., (1988): p. 460.

Note: The status of Peltaria angustifolia, in the Flora Palaestina area, is discussed by Danin (1999: p.158).

Pennisetum clandestinum Chiov.

Note: Although not reported before for Israel this plant is rather common in disturbed habitats, mainly in ditches along roads where the roadside management is practiced with herbicides and in heaps of remains of house construction. In my database it is listed from 25 squares of 5X5 km mainly in the Mediterranean territory of Israel . The record of this species from the Judean Desert is from the banks of Nahal Qidron (Wadi Nar), at present polluted and became an opened sewage canal which streams the year round.

Phagnalon nitidum Fresen.

Note: The first specimen in HUJ is: Jordan, Jebel Umm Adami, 55 km ESE of Aqaba,  a rocky wadi, 10.5.1998, Danin 980213.

Phagnalon sinaicum Bornm. & Kneuck.

Note: The first specimen in HUJ is: Jordan, Jebel Umm Adami, 55 km ESE of Aqaba,  a rocky wadi, 10.5.1998, Danin 980301.

Phlomis chrysophylla Boiss.

Ref.: Danin, in Greuter & Raus (1982b): p. 198.

Phyllanthus rotundifolius Klein ex Willd.

Note: The first specimen in HUJ is: Judean Mountains , Ramot, in a garden, 26.10.1978, H. Almagor, det. D. Heller.

Physalis angulata L.

Note: The first specimens in HUJ: Esderaelon Plain, Nahalal, a weed in a garden, 10.3.1977, A. Dafni, det. W.G. D’Arcy.

Picris babylonica Hand.-Mazz.

Note: A few specimens previously determined as Picris cyanocarpa Boiss.  in HUJ were renamed in 1999 as P. babylonica by M. Sualla (B) who studies this group from the Middle East .

Pinus brutia Ten.

Note: A tree of Mediterranean countries north and west of Israel , introduced for afforestation during the 20th century. It has casual occurrences near areas planted with P. brutia. Hybrids with Pinus halepensis occur occasionally throughout the Mediterranean part of the country. Pinus  halepensis is the pine species most frequently planted in the Mediterranean mountains of Israel . A few spontaneous small populations of the latter occur in the Judean and Samarian Mountains . Large populations of it occur in Mt. Carmel and the Upper Galilee . Both pine species establish themselves in areas where dwarf-shrub and shrub communities cover the area. During the processes of plant succession the dwarf-shrubs discard the annual associations typical to the early stages of plant succession on old fields (Danin 1995). Casual trees of Pinus brutia and hybrids Pinus halepensis X brutia occur among trees of P. halepensis in such shrub and semishrub associations. It also occurs in roadsides on fissured limestone outcrops where hills were cut during the preparation of the road and planted forests occur in the vicinity.

Plantago sinaica (Barnéoud) Decne.

Specimens seen: Edom , Dana Reserve, Wadi Barra area, below the forest-ranger station, 2 km SE of the visitor center, in crevices and near smooth-faced white sandstone outcrops,  14.5.1996, Danin 963020.

Ref.: Danin (1997) 27: p. 172.

Note: This is a common component of the rock vegetation of the smooth-faced granite at the high elevation belt of S Sinai . In SW Jordan only a few specimens were found so far and only at the collection site of the above specimen.

Polygonum argyrocoleum Steud. ex G.Kunze

Ref:Danin, in Greuter & Raus (1989): p. 41.

Polygonum aviculare L.

Ref:Danin, in Greuter & Raus (1989): p. 41.

Potentilla supina L.

Note: First specimen in HUJ: Negev Highlands, near Yeroham, wet soil in banks of the (artificial) lake, 15.7.1971, Danin.

Pteridium aquilinum (L.) Kuhn

Ref.: Danin, in Greuter (1980): p. 21.

Ranunculus lateriflorus DC.

Note:Danin, in Greuter & Raus (1989): p. 46.

Robinia pseudoacacia L.

Note: An adventive tree of N American origin, planted as an ornamental tree. Occasionally germinates and establishes itself in disturbed ground of urban areas and roadside in the Mediterranean territories of the study area.

Rorippa prostrata (J.P.Bergeret) Schinz & Thell.

Note: An adventive plant, which invaded Israel during the late 1980’s by roots attached to bulbs which were introduced from The Netherlands (Feinbrun-Dothan & Danin 1991). Multiplies mainly vegetatively. The present status and distribution is not clear.

Sageretia thea (Osbeck) M.C.Johnst.

Note:Künne in Greuter & Raus (1989): p. 48.

Sagina maritima G.Don

Ref.: Danin (1992): p. 178.

Salsola cyclophylla Baker

Note: Danin (1973): p. 22. 

Salsola gaetula (Maire) Botsch.

Ref.: Danin, in Greuter & Raus (1982a): p. 40.

Salsola orientalis S.G.Gmel.

Ref.: Danin, in Greuter & Raus (1982a): p. 40.

Salvinia natans (L.) All.

Note: The first specimen in HUJ: Sharon, Netanya, Dora’s Pool, 5.9.1988.

Sambucus ebulus L.

Note: Liston, in Greuter & Raus (1987): p. 439.

Sambucus nigra L.

Note: First specimen in HUJ: Upper Galilee , Ein Zeitim near Safad, wadi, 23.6.1925, Naftolsky.

Scandix australis L.

Note: Danin, in Greuter & Raus (1989): p. 48.

Scandix grandiflora L.

Note: Danin, in Greuter & Raus (1989): p. 48.

Schinus molle L.

Ref.: Danin, in Greuter (1981): p. 29.

Note: Introduced as an ornamental tree from S. America , Andes . Although being used as an ornamental plant much more than Schinus terebinthifolis, S. molle is much less common as growing independently in places where it was not actually planted. It is found casually in roadsides and wastelands.

Schinus terebinthifolius Raddi

Ref.: Danin, in Greuter & Raus (1989): p. 28.

Note: A tree that was introduced from Brazil to Europe as an ornamental plant during the 19th century. It is not clear when it was introduced to Israel . It has a constant occurrence in human-managed habitats in lowlands but in small quantities. The kind of habitats S. terebinthifolia occupies is similar to those of Lantana camara, but the frequency is lower. It is found in lowlands in almost abandoned and abandoned citrus groves, trickle-pipe-irrigated date palm plantations, and in wastelands. It has small red fruits and is likely to being dispersed by small birds through endozoochory.

Schoenefeldia gracilis Kunth

Note: First specimen in HUJ: Pleshet, Kvouzat Schiler, a weed in irrigated lawn, 5.12.1998, B. Rubin, det. Danin & Scholz.

Scleranthus orientalis Roessler

Note: The first specimen in HUJ: Golan, Mas’ada forest, among basalt rocks, 4/1973, Shmida.

Secale cereale L.

Note: Found casually at the herbicides-sprayed roadsides of the Mediterranean territories. The origin of the specimens may be from grains transferred for various reasons in these highways. The individuals in the roadsides do not tend to grow in-groups and may indicate new contamination annually.

Seidlitzia cinerea (Moq.) Bge. ex Botsch.

Note: The first and only specimen in HUJ: Edom , environs of Aneze, 22.7.1942, Zohary & Feinbrun, det. Botschantzev.

Senna obtusifolia (L.) Irwin & Barneby

Ref.: Joel &  Liston (1986).

Sesbania sesban (L.) Merr.

Ref.: Dafni & Heller (1990).

Setaria verticillata (L.) P. Beauv.

Specimens seen: Israel : Kiryat Anavim, 6.vii.1996, Danin, Jerusalem . Jordan , Edom : 1 km W of Adhruh, Irrigated fruit trees plantation, 26.x.1997, Danin 972704 (HUJ).

Ref.: Danin & Scholz (1997): pp. 177-179.

Note: The two taxa in the Setaria verticillata complex are clearly recognized in Israel and Jordan . There is no transition in the diagnostic morphological characters of the common Setaria adhaerens (Forssk.) Chiov., and the newly introduced and relatively rare S. verticillata.

Sida acuta Burm.f.

Note: First specimen at HUJ: Beit Shean Valley , Tirat Zevi, near fishponds, 29.6.1980, Dafni.

Siebera nana (DC.) Bornm.

Ref.: Baierle et al., (1988).

Silene libanotica Boiss.

Note: A specimen of this species, collected by E. Boissier in Hebron is deposited at Boissier Herbarium in Geneve (!).

Solanum dulcamara L.

Note: The first specimen in HUJ is ex herbarium Postianum: Plantae Giladenses, 4.5.1886 (the exact location is not clear). At present it is a rather common plant in the spring vegetation of the Banyas springs at the meeting zone of the Golan, the Hermon, and the Hula-Dan Valleys .

Solanum laciniatum Aiton

Note: The first specimens in HUJ: Golan, Chispin, drainage channel, escaped from cultivation, 15.5.1985, O. Cohen. At the same year R. Ducas collected it from a few locations with disturbed ground.

Sonchus microcephalus Mejias

Ref.: Danin (1992): p.193.

Sterigmostemum sulphureum (Banks & Sol.) Bornm.

Note: Al-Eisawi (1985b): p. 953.

Stipa ehrenbergiana Trin. & Rupr.

Ref.: Freitag (1985).

Stipa holosericea Trin.

Ref.: Freitag (1985).

Stipagrostis drarii (Täckh.) de Winter

Ref.: Baierle et al., (1988): p. 465; Danin (1994): pp. 59‑61

Stipagrostis uniplumis (Licht.) de Winter

Ref.: Al-Eisawi (1985a).

Tagetes minuta L.

Ref.: Dafni & Heller (1990).

Teesdalia coronopifolia (J.P.Bergeret) Thell.

Note: The first record in HUJ is: Golan: nr Masa’ada, shade among oaks, 7.4.1987, Danin.

Thalictrum isopyroides C.A.Mey.

Ref.: Boulos et al., (1975): p. 369 (for Jordan ), Danin, in Greuter & Raus (1984a): p. 54 (for Israel ).


Note: This rare steppe plant was observed and collected in the Negev Highlands a few more times.

Torilis webbii S.L.Jury

Ref: Jury (1987): p. 297.

Trianthema portulacastrum L.

Note: Danin (1981): p. 27.

Trifolium glomeratum L.

Note: Fragman, in Greuter & Raus (1999): p. 60. This Trifolium which was not mentioned by Zohary (1972) was well known to M. Zohary from the Golan. Fragman (1999) reports the first occurrence of this plant from the Upper Galilee . In my database it is recorded from 7 squares of 5x5 km, mainly from volcanic ash at the vicinity of volcanic cones of the eastern Golan.

Trifolium hirtum All.

Note: The first and only specimen in HUJ: Golan, Bab el Hawa, 31.3.1969, Dafni, det. D. Heller.

Trifolium sylvaticum Gérard

Note: Golan, near Quneitra, tuff quarry, 25.4.1969, Shmida, det. D. Heller.

Trigonella lunata Boiss.

Ref.: Boulos & Al-Eisawi (1977b).

Trigonella sibthorpii Boiss.

Ref.: Danin & Small (1989).

Note: The only population of this species found so far in Israel developed at a roadside in the Judean Desert Spring 1974. In the last few years, as a result of expanding the road width, the site of Trigonella sibthorpii population disappeared                                                      

Trigonella spruneriana Boiss.

Note: The first and only specimen in HUJ: Golan, near Susita, chalk, 8.3.1984, Liston 7-84-176/18.

Triticum urartu Tumanian ex Gandilyan

Ref.: Valkoun et al., (1998).

Typha angustifolia L.

Ref.: Danin, in Greuter & Raus (1989): p. 48.

Note: In addition to the populations found in the newly exposed coastal area of the Kinneret in drought years and depressions with high underground water table near Rishon LeZion, it was found recently near fresh water springs at Ain al Fidan , Edom (SW Jordan ), 17.7. 2000, Danin, 2K0301(HUJ).

Urochloa panicoides P.Beauv.

Note: The first specimen in HUJ: Pleshet: 3 km S of Rehovot, Kibbutz Givat Brener, a ditch draining fields near an asphalt road, 4.x.1998, Danin.

Urochloa texana (Buckley) R.Webster

Note: The first specimen in HUJ: Northern Negev : Ofaqim, a weed in irrigated flowerbed. 21.x.1989, Danin 38781.

Ref.: Danin & Cope, in Greuter & Raus (1995): p. 175; first reported as Brachiaria texana (Buckley) S.T. Blake. This plant was collected again and was observed to be a common weed in a cotton field in the Philistean Plain and was re-determined by Prof. H. Scholz.

Veronica panormitana Tineo subsp. baradostensis (M.A.Fisch.) M.A.Fisch.

Specimen seen: Jordan , Gilead : Zubia Reserve, north of Ajloun, maquis of Quercus calliprinos, alt. 1,000 m. above sea level, 16.3.1996, Danin 960845, det. M.A. Fischer.

Note: Danin (1997): p. 172. After the determination of this diploid taxon with small petals by the expert, Prof. Dr. M.A. Fischer and its distinction from the common polyploid V. cymbalaria Bodard, which has larger petals,  it was collected also from the following districts: Sharon Plain , Samaria , Shefela, and Judean Mts.

Veronica polifolia Benth.

Specimens seen: Edom, northern section of Dana Reserve, the Cupressus sempervirens reserve (4 km S of Bseara),  1500 m, sandy soil, 15.5.1996, Danin 963101.

Ref.: Danin (1997): p. 173.

Note: This semi-shrub, common in depressions of the subalpine Mt. Hermon (dolines) grows in a unique semi-shrub community in the steppe-forest area on sandy soil.

Vicia benghalensis M.Bieb.

Note: The first specimen in HUJ: Hula Valley , Amir, 1.4.1941, F. Weisman.

Vicia lathyroides L.

Note: First specimen in HUJ: Golan, Masa’ada forest, 20.4.1969, Danin.

Vincetoxicum dionysiense Mouterde

Note: First specimen in HUJ:  Golan, Nahal Yehudiya 3 km ENE of Katzrin, 4.9.1985, Kaplan, det. A. Liston.

Vitis vinifera L. subsp. sylvestris (C.C.Gmel.) Hegi

Note: Rottenberg (1998): p.142 presents the summary of his observations in spontaneous populations of this taxon, and concludes that it occurs as a component in riparian vegetation of the sources of the Jordan River .

Vulpia membranacea (L.) Dumort

Note: The first specimen in HUJ: Sharon, Magdiel, near Tel Aviv, sandy field, 27.4.1928, Eig, det. H. Scholz 2000. The specimen we have in HUJ is a photocopy of the specimen in B (in Berlin ). It was sent from our institute many years ago as part of “Flora Palaestinae Exicata”, No. 16 Vulpia uniglumis Dum.

Vulpia persica Boiss.

Ref.: Danin, in Greuter & Raus (1987): p. 452. Danin & Scholz (1994): pp. 257-259.

Washingtonia filifera Wendl.

Note: Indigenous tree confined to fresh water springs in SW North America (California and Arizona ). Introduced as an ornamental plant from California at the beginning of the 20th century. Establishing itself in disturbed ground with low competition and sufficiently wet ground. These conditions may be found in abandoned or almost-abandoned citrus groves of the coastal plain, in gardens at the Mediterranean territories, in roadsides and in trickle-pipe-irrigated date-palm groves. Prevailing spontaneously near desert springs in California and Arizona at a high range of elevations, W. filifera is expected to grow at a wide range of elevations in Israel as well.


3. List of species which were mis-identified in Flora Palaestina

 

There are a few species in Flora Palaestina, which, according to the contemporary opinions expressed in the quoted literature, were mis-identified. The erroneous name is presented in Italics and between inverted commas. The correct name is typed in boldface.

 

 Allium stamineum“. Allium daninianum Brullo, Pavone & Salmeri, Ref: Willdenowia 26: 237-244 (1996).

Note: See discussion above under Allium daninianum.

Arabis nova”. Arabis auriculata Lam

Ref: Greuter et al., (1986): p. 53.

 Aristolochia maurorum  Aristolochia bottae Jaub. & Spach

 Ref: Greuter et al., (1984): p. 53.

Artemisia herba-alba”  Artemisia sieberi Besser

 Ref: Leonard (1984): p. 20.

Asphodelus aestivusAsphodelus ramosus L

 Ref: Diaz Lifante & Valdes, in Danin (1992): p. 61.

Atriplex hastata”. Atriplex prostrata Boucher ex DC.

 Ref: Greuter et al., (1984): p. 293.

Brachypodium pinnatum”  Brachypodium sylvaticum (Huds.) P.Beauv.

 Note: Thanks are due to Prof. H.Scholz for his comments on the need to re-determine this species.

Bromus brachystachys”  Bromus pseudobrachystachys H.Scholz

Note: In fact most of the specimens of Bromus determined as “Bromus brachystachys” by Prof. N. Feinbrun (in HUJ) and included in Flora Palaestina IV (Feinbrun-Dothan 1986) were determined by Prof. H.Scholz (summer 2000) as Bromus tigridis Boiss. & Noae. Only a few specimens of Bromus in HUJ have been named B. pseudobrachystachys by an expert. The whole genus in Flora Palaestina area deserves revision.

Capparis ovata”. Capparis sicula Veill.

  Ref.: Danin, in Greuter 1981: p. 41; Greuter et al., (1984): p. 155.

Chaenorrhinum persicum”. Hueblia calycina (Banks & Sol.) Speta

 Cleome arabica”. Cleome amblyocarpa Barratte & Murb

 Ref: Greuter et al., (1984): p. 155.

Colchicum bowlesianum”. Colchicum feinbruniae Persson

Commicarpus verticillatus”. Commicarpus helenae (J.A.Schultes) Meikle

 Ref: Greuter et al., (1989): p. 244.

Corrigiola litoralis subsp. telephiifoliaCorrigiola palaestina Chaudhri

 Ref: Greuter et al., (1984): p. 185.

Corynephorus divaricatus” Corynephorus articulatus (Desf.) P.Beauv.

 Ref: According to Prof. H. Scholz (pers. comm.) C. divaricatus  is a west Mediterranean species.

Cymbopogon parkeri” Cymbopogon commutatus (Steud.) Stapf

 Cynosurus elegansCynosurus effusus Link

Cyperus conglomeratus”. Cyperus macrorrhizus Nees

  Ref:   Danin & Kukkonen (1995).

Cytinus hypocistis(L.) L.   Cytinus ruber (Fourr.) Kom

Note: The Cytinus root-parasite of the genus Cistus, which grows in Israel, has flowers with white corolla and bracts with red colour. It fits the diagnostic characters of C. ruber and not of C. hypocistis.

 Erodium subtrilobum”   Erodium neuradifolium Delile

 Ref:  Greuter et al., (1986): p. 255.

Himantoglossum affine” . Himantoglossum caprinum (M.Bieb.) Spreng.

 Note: Prof. A. Dafni (pers. comm.) communicated the nomenclature changes.

 Hypecoum imberbe”. Hypecoum dimidiatum Delile

  Ref:   Greuter et al., (1989): p. 281.

 Lathyrus digitatusLathyrus spathulatus Čelak.

 Ref:  Greuter et al., (1989): p. 117.

 Limonium meyeri”. Limonium narbonense Mill

 Ref:  Greuter et al., (1989): p. 344.

 Limonium oleifolium”. Limonium virgatum (Willd.) Fourr

  Ref: Greuter et al., (1989): p. 343.

Ononis reclinata”. Ononis mollis Savi

Ref: Valdes in Danin (1992): p. 152.

Note: Following Valdes’ comment the material of the two taxa was studied in HUJ. I found there only O. mollis.

Orchis laxiflora” . Orchis dinsmorei (Schlechter) H.Baumann & Dafni

Note: Prof. A. Dafni (pers. comm.) communicated the nomenclature changes.

Orchis papilonacea” Orchis caspia Trautv.

Note: Prof. A. Dafni (pers. comm.) communicated the nomenclature changes.

 Pentatropis spiralis”. Pentatropis nivalis (J.F.Gmel.) Field & Wood

 Ref: Greuter et al., (1984): p. 57.

Petrorhagia cretica”. Petrorhagia zoharyana A.Liston

 Polygala sinaica”.  Polygala negevensis Danin

 Ref: Danin (1987).

Polygonum patulum”  Polygonum bellardii All.

 Ref: Greuter et al., (1989): p. 358.

Raphanus aucheri”. Quidproquo confusum Greuter & Burdet

Ref: Greuter & Burdet, in Greuter  & Raus (1983): p. 94; Greuter et al., (1986): p. 149.

“Reboudia pinnataErucaria microcarpa Boiss

 Ref: Greuter et al., (1986): p. 104.

Salsola longifolia“. Salsola oppositifolia Desf

Ref: Greuter et al., (1984): p. 310.

 Salsola tetragona”. Salsola cyclophylla Baker

  Ref: Danin (1973): p. 22.

 Sedum pallidum” Sedum rubens L

 Ref:  H.T. Hart, in Danin (1992): p.187.

“Serapias vomeracea” Serapias levantina H.Baumann & Künkele

Ref: Prof. A. Dafni (pers. comm.) communicated the nomenclature changes.

Setaria glauca” Setaria pumila (Poir.) Schult.

 Note: Setaria glauca”. Setaria glauca (L.) P.Beauv. is not a taxonomic synonym of  S. pumila. It was erroneously applied by many authors to S. pumila, however, the Linnaean basionym Panicum glaucum L. is known at present as the cultivated millet Pennisetum glaucum (L.) R.Br.

 Setaria verticillata  Setaria adhaerens (Forssk.) Chiov

 Ref: Danin & Scholz (1997).

Spergularia media”. Spergularia maritima (All.) Chiov

 Ref: Greuter et al., (1984): p. 283.

Stipa barbata”  Stipa arabica Trin. & Rupr.

 Ref: Freitag (1985): p. 461.

Stipa lagascae” Stipa pellita (Trin. & Rupr.) Tzvelev 

Ref:  H. Scholz (pers. comm.).

Note: A few specimens labeled Stipa lagascae in HUJ were determined Stipa pellita and others as Stipa holosericea by Prof. Freitag and by Prof. Scholz.

 

4. List of species the names of which in Flora Palaestina are regarded at present as synonyms

In the following section the correct updated names are presented in boldface; the synonyms used in Flora Palaestina are written in Italics following the abbreviation Fl.Pal. Most changes follow the conclusions of taxonomic investigations presented in the 3 volumes of MedChecklist (Greuter et al. 1984, 1986, and 1989).

 

Acacia pachyceras O. Schwartz var. najdensis (Chaudhary) Boulos         

Fl.Pal.: Acacia gerrardii Benth. subsp. negevensis Zohary

Notes: In his account of the Acacia species of the Arabian Peninsula, Boulos (1995: 335-336) disconnects the geographical distribution-relations of the species of Acacia, known as TALKH in several Arab speaking countries (e.g. Sinai, Jordan, and Arabia) from the African Acacia gerrardii Benth. Contrary to Zohary (1972), he ties it to the species described from Yemen by Schwartz (1939): “In Arabien im innerjemenitischejhn hochland”. Being acquainted from the field with the “Talkh” in Israel, Sinai and Jordan, and studying both this species and Acacia gerrardii in the herbarium (at Kew) I agree with Boulos (1995). I think as Boulos did that the Near Eastern Talkh of Flora Palaestina should be named Acacia pachyceras O.Schwartz var najdensis (Chadhry) Boulos. The comment of Chaudhary (1998) disregards the fact that Schwartz (1939) described Acacia pachyceras from Yemen. Boulos (1995) thus selected a neotype for this species (Yemen: J.R.I. Wood 2401 [K!]). One of the best field characters that helps recognizing Acacia pachyceras is it easily pilling bark into white-cream longitudinal strips. Schwartz gives it in his diagnosis of A. pachyceras “tecti primo pilis albis brevibus intermitis longioribus”. Several collectors of Acacia gerrardii Benth. in the Sudan and in Ethiopia (in K !) write that tree’s bark is black and deeply longitudinally fissured. This property is never found in A. pachyceras of Israel, Jordan, and Sinai.

Acer obtusifolium Sm; Fl.Pal.: Acer syriacum Boiss. & Gaill. [syn.]

Achillea arabica Kotschy; Fl.Pal.: Achillea biebersteinii Afan.

Ref:  Valant-Vetschera (1999): p.144.

Adonis microcarpa DC; Fl.Pal.: Adonis cupaniana Guss. [syn.]

Adonis palaestina Boiss.; Fl.Pal.: Adonis aestivalis L. var. palaestina (Boiss.) Zohary [syn.]

Aegilops vavilovii (Zhuk.) Chennav.; Fl.Pal.: Aegilops crassa Boiss. subsp. vavilovii Zhuk. [syn.]

Aerva javanica (Burm.f.) Juss. ex Schult; Fl.Pal.: Aerva persica (Burm.f.) Merr. [syn.]

Aetheorhiza bulbosa (L.) Cass.; Fl.Pal.: Crepis bulbosa (L.) Tausch. [syn.]

Agathophora alopecuroides (Delile) Fenzl ex Bunge; Fl.Pal.: Halogeton alopecuroides (Del.) Moq. [syn.]

Ajuga chamaepitys (L.) Schreb. subsp. chia (Schreb.) Arcang.; Fl.Pal.: Ajuga chia Schreb. [syn.]

Alhagi graecorum Boiss.; Fl.Pal.: Alhagi maurorum Medik. [syn.]

Allium neapolitanum Cirillo; Fl.Pal.: Allium neapolitanum Cyr.

Allium trifoliatum Cirillo; Fl.Pal.: Allium trifoliatum Cyr.

Alyssum aureum (Fenzl) Boiss.; Fl.Pal.: Alyssum meniocoides Boiss.

Alyssum simplex Rudolphi; Fl.Pal.: Alyssum minus (L.) Rotm. var. minus [syn.]

Alyssum strigosum Banks & Sol.; Fl.Pal.: Alyssum minus (L.) Rothm. var. strigosum (Banks & Sol.) Stoj. [syn.]

Amaranthus cruentus L.; Fl.Pal.: Amaranthus hybridus L. [syn.]

Amaranthus muricatus Gillies ex Hicke; Fl.Pal.: Amaranthus muricatus Moq. [syn.]

Amaranthus viridis L.; Fl.Pal.: Amaranthus gracilis Desf. [syn.]

Ammannia baccifera L.; Fl.Pal.: Ammania aegyptiaca Willd. [syn.]

Anchusa azurea Mill.; Fl.Pal.: Anchusa italica Retz. [syn.]

Arabis alpina L. subsp. caucasica (Willd.) Briq.; Fl.Pal.: Arabis caucasica Schltdl. [syn.]

Aristida adscensionis L.; Fl.Pal.: Including Aristida coerulescens Desf.

Arthrocnemum macrostachyum (Moric.) K.Koc; Fl.Pal.: Arthrocnemum macrostachyum (Moric.) Moris & Delponte [syn.]

Arundo pliniana Turr; Fl.Pal.: Arundo plinii Turra

Asparagus horridus L.; Fl.Pal.: Asparagus stipularis Forssk. [syn.]

Asplenium ceterach L.; Fl.Pal.: Ceterach officinarum DC. [syn.]

Asplenium sagittatum (DC.) A.J.Bange; Fl.Pal.: Phyllitis sagitata (DC.) Guinea & Heywood [syn.]

Asplenium onopteris L.; Fl.Pal.: Asplenium adiantum-nigrum L. subsp. onopteris (L.) Heufl. [syn.]

Asteriscus hierochunticus (Michon) Wiklund; Fl.Pal.: Asteriscus pygmaeus (DC.) Coss. & Dur. [syn.]

Astomaea seselifolium DC. Fl.Pal.: Astoma seselifolium DC.

Astragalus aleppicus Boiss.; Fl.Pal.: Astragalus giladensis Eig [syn.]; A. postii Eig [syn.]; A. galilaeus Freyn & Bornm. [syn.]; A. feinbruniae Eig ex Rech.f. [syn.]

Astragalus arnilobus Kar. & Kir.; Fl.Pal.: Astragalus gyzensis Delile ex Bge. [syn.]; A. hauarensis Boiss. [syn.]

Astragalus asterias Steve; Fl.Pal.: Astragalus cruciatus Link [syn.]

Astragalus campylorrhynchus Fisch. & C.A.Mey.; Fl.Pal.: Astragalus negevensis Zohary & Fertig  [syn.]

Astragalus caprinus L. subsp. lanigerus (Desf.) Maire ; Fl.Pal.: Astragalus platyraphis Fisch. exBge. [syn.]; A. beershabensis Eig & Sam. [syn.], A. alexandrinus Boiss. [syn.]

Astragalus crenatus Schultes; Fl.Pal.: Astragalus corrugatus Bertol. [syn.]

Astragalus dactylocarpus Boiss. subsp. acinaciferus (Boiss.) E.Ot; Fl.Pal.: Astragalus acinaciferus Boiss. [syn.]

Astragalus kotschyanus Boiss.; Fl.Pal.: Astragalus zemeraniensis Eig [syn.]

Astragalus pelecinus (L.) Barneb; Fl.Pal.: Biserrula pelecinus L. [syn.]

Atriplex glauca L.; Fl.Pal.: Atriplex stylosa Viv. [syn.]

Atriplex portulacoides L.; Fl.Pal.: Halimione portulacoides (L.) Aellen [syn.]

Atriplex prostrata Boucher ex DC; Fl.Pal.: “Atriplex hastata”.

Atriplex sagittata Borkh.; Fl.Pal.: Atriplex nitens Schkuhr [syn.]

Bassia arabica (Boiss.) Maire & Weille; Fl.Pal.: Chenolea arabica Boiss. [syn.]

Bassia indica (Wight) A.J.Scott; Fl.Pal.: Kochia indica Wight [syn.]

Bellis sylvestris Cirillo; Fl.Pal.: Bellis silvestris Cyrill.

Beckmannia eruciformis (L.) Host; Fl.Pal.: Beckmannia erucaeformis (L.) Host

Note: According to the contemporary nomenclatural rules, an epithet composed of two words can not be joined by a diphthong (T. Cope, 8/2000, pers. comm.).

Bilacunaria boissieri (Reut. & Huasskn.) Pimenov & Tichomiro; Fl.Pal.: Hippomarathrum boissieri Reut. & Hausskn. [syn.]

Bituminaria bituminosa (L.) C.H.Stirt.; Fl.Pal.: Psoralea bituminosa L. [syn.]

Bituminaria flaccida (Nábelek) Greute; Fl.Pal.: Psoralea flaccida Nábelek [syn.]

Blepharis in Israel

Note: According to K. Vollesen (pers. comm., 8/2000), who compiled a monograph of the genus Blepharis, the common species of Israel, Jordan and Sinai, is not B. ciliaris (L.) B.L.Burtt (sensu Feinbrun-Dothan 1978: p. 218). Vollesen indicated that B. ciliaris is actually an endemic species of Iran-Afghanistan. The populations of Blepharis attenuata Napper, as it is known in the phytosociological research in Israel, are confined to semisteppe batha (Danin & Solomeshch 1999). It is a diagnostic species of the Balloto-Sarcopoterion spinosi Danin & Solomeshch, an alliance in the class Ballotetea undulatae Danin & Solomeshch. When Napper (1972) described this species it was considered as a mesophitic plant and was later declared as a diagnostic of a marginal Mediterranean plant community. It was considered as an independent species differing from the other, more drought and heat resistant taxon of the warm desert areas of the Dead Sea, Arava Valley, and the coastal area of eastern Sinai (B. ciliaris sensu Feinbrun 1978). However, according to the present revision of Vollesen (2000, in press), B. attenuata is the right name for both taxa, as so far there is no constant diagnostic character which may be used to distinguish between the two ecological and geographical different races.

Brachypodiumdistachyum (L.) P.Beauv.; Fl.Pal.: “Brachypodium distachyon”

Note: The spelling “distachyon” by Beauvoix was inaccurate (T. Cope 8/2000, pers. comm.).

Bunium paucifolium DC; Fl.Pal.: Bunium elegans (Fenzl) Freyn [syn.]

Bupleurum orientale Snogerup; Fl.Pal.: Bupleurum tenuissimum L. [syn.]

Callipeltis factorovskyi (Eig) Ehrend.; Fl.Pal.: Waburgina factorovskyi Eig [syn.]

Callitriche brutia Petagn; Fl.Pal.: Callitriche pedunculata DC. [syn.]

Callitriche lenisulca Clavaud; Fl.Pal.: Callitriche palustris L  [syn.].

Callitriche truncata Guss; Fl.Pal.: Callitriche hermaphroditica L. [syn.]

Capparis aegyptia Lam; Fl.Pal.: Capparis spinosa L. var. aravensis Zohary [syn.]

Capparis sinaica Veill.; Fl.Pal.: Capparis cartilaginea Decne. [syn.]

Carduus acicularis Bertol.; Fl.Pal.: Carduus argentatus L. var. esdraelonicus (Boiss.) Boiss. [syn.]

Carex guestphalica (Boenn. ex Rchb.) Boenn. ex O.Lang; Fl.Pal.: Carex divulsa Stocks subsp. leersii (Kneuck.) W.Koch [syn.]

Carlina libanotica Boiss. subsp. microcephala (Post) Meusel & Dittrich; Fl.Pal.: Carlina hispanica Lam. subsp. galilaea Meusel & Kästner [syn.]

Centaurea pumilio L.; Fl.Pal.: Aegialophila pumilio (L.) Boiss. [syn.]

Note: T. Raus (pers. comm.) suggests returning it to Centaurea because there are many sections in Centaurea and in Flora Europaea 4: 269 the authors returned all back to Centaurea. Feinbrun-Dothan (1978) took one small group and retained Aegialophila.

Centropodia forskalii (Vahl) Cope; Fl.Pal.: Asthenatherum forskalii (Vahl) Nevski [syn.]

Cerastium glomeratum Thuill.; Fl.Pal.: Cerastium viscosum L. [syn.]

Cerasus microcarpa (C.A.Mey.) Boiss.; Fl.Pal.: Cerasus microcarpa (C.A.Mey.) C.Koch [syn.]

Ceratocapnos turbinata (DC.) Lide; Fl.Pal.: Ceratocapnos palaestinus Boiss. [syn.]

Cheilanthes acrostica (Balb.) Tod.; Fl.Pal.: Cheilanthes fragrans (L.) Webb & Berth. [syn.]

Chiliadenus iphionoides (Boiss. & Blanche) Brullo; Fl.Pal.: Varthemia iphionoides Boiss & Blanche [syn.]

Chiliadenus montanus (Vahl) Brullo; Fl.Pal.: Varthemia montana (Vahl) Boiss. [syn.]

Chrozophora tinctoria (L.) Raf.; Fl.Pal.: Chrozophora tinctoria (L.) Ad.Juss. [syn.]

Cicer judaicum Boiss.; Fl.Pal.: Cicer pinnatifidum Jaub. & Spach [syn.]

Cichorium endivia L. subsp. divaricatum (Schousb.) P.D.Sell; Fl.Pal.: Cichorium pumilum Jacq. [syn.]

Cleome arabica L.; Fl.Pal.: Cleome trinervia Fresen. [syn.]

Colchicum troodi Kotsch.; Fl.Pal.: Colchicum decaisnei Boiss. [syn.]

Commicarpus plumbagineus (Cav.) Standle; Fl.Pal.: Commicarpus africanus (Lour.) Dandy [syn.]

Consolida incana (E.D.Clarke) Munz; Fl.Pal.: Consolida rigida (DC.) Bornm. [syn.]

Convolvulus auricomus (A.Rich.) Bhandari; Fl.Pal.: Convolvulus glomeratus Choisy [syn.]

Conyza stricta Willd.; Fl.Pal.: Conyza triloba Decne. [syn.]

Cosentinia vellea (Aiton) Tod.; Fl.Pal.: Cheilanthes catanensis (Cosent.) H.P. Fuchs [syn.]

Crucianella aegyptiaca L.; Fl.Pal.: Crucianella herbacea Forssk. [syn.].

Ref.: C. aegyptiaca is mentioned in Hepper & Friis (1994) as the legitimate name, C. herbacea is therefore a synonym.

Cucumis acidus Jacq.f.; Fl.Pal.: Cucumus callosus (Rottler) Cogn. [syn.]

Cyperus michelianus (L.) Delile subsp. pygmaeus (Rottb.) Asch. & Graebn.; Fl.Pal.: Cyperus pygmaeus Rottb. [syn.]

Daucus broteri Ten.; Fl.Pal.: Daucus bicolor Sm. [syn.]

Daucus durieua Lang; Fl.Pal.: Daucus subsessilis Boiss. [syn.]

Daucus glaber (Forssk.) Thell.; Fl.Pal.: Daucus litoralis Sm. [syn.]

Desmazeria philistaea (Boiss.) H.Scholz; Fl.Pal.: Cutandia philistaea (Boiss.) Jackson  [syn.]

Deverra tortuosa (Desf.) DC.; Fl.Pal.: Pituranthos tortuosus (Desf.) Asch. & Schweinf. [syn.]

Deverra triradiata Hochst. ex Boiss.; Fl.Pal.: Pituranthos triradiatus (Hochst. ex Boiss.) Asch. & Schweinf. [syn.]

Dianthus monadelphus Vent. subsp. judaicus (Boiss.) Greuter & Burdet; Fl.Pal.: Dianthus judaicus Boiss. [syn.]

Dichanthium foveolatum (Delile) Robert; Fl.Pal.: Eremopogon foveolatus (Delile) Stapf [syn.]

Dittrichia graveolens (L.) Greuter; Fl.Pal.: Inula graveolens (L.) Desf. [syn.]

Dittrichia viscosa (L.) Greuter; Fl.Pal.: Inula viscosa (L.) Aiton [syn.]

Doellia bovei (DC.) Anderb.; Fl.Pal.: Blumea bovei (DC.) Vatke [syn.]

Ref:  Anderberg (1995).

Dorycnium hirsutum (L.) Ser.; Fl.Pal.: Bonjeana hirsuta (L.) Rchb. [syn.]

Dorycnium rectum (L.) Ser; Fl.Pal.: Bonjeana recta (L.)Rchb. [syn.]

Dryopteris pallida (Borry) C.Chr. ex Maire & Petitm. subsp. libanotica (Rosenst.) Nardi; Fl.Pal.:  Dryopteris villarii (Bellardi) H.Woynar ex Schniz & Thell. [syn. p.p.]

Eclipta prostrata(L.) L.; Fl.Pal.: Eclipta alba (L.) Hassk.

Eigia longistyla (Eig) Soják; Fl.Pal.: Stigmatella longistyla Eig [syn.]

Elytrigia elongata (Host) Nevski; Fl.Pal.: Elymus elongatus (Host) Runemark [syn.]

Elytrigia intermedia (Host) Nevski; Fl.Pal.: Elymus hispidus (Opiz) Meld. [syn.]

Elytrigia juncea (L) Nevski; Fl.Pal.: Elymus farctus(Viv.) Runemark ex Melderis [syn.]

Enneapogon desvauxii P.Beauv.; Fl.Pal.: Enneapogon brachystachyus (Jaub. & Spach) Stapf [syn.]

Ephedra aphylla Forssk.; Fl.Pal.: Ephedra alte C.A.Mey. [syn]

Ref.: Danin & Hedge (1973): pp. 264-265.

Ephedra ciliata Fisch. & C.A.Mey.; Fl.Pal.: Ephedra peduncularis Boiss. [syn.]

Ref.: Freitag & Maier-Stolte (1994).

Ephedra foeminea Forssk.; Fl.Pal.: Ephedra campylopoda C.A.Mey. [syn.]

Ref.: Freitag & Maier-Stolte (1994).

Epilobium tetragonum L. subsp. tournefortii (Michalet) Léveille; Fl.Pal.: Epilobium tournefortii Michalet  [syn.]

Eragrostis cilianensis (All.) Janch; Fl.Pal.: Eragrostis cilianensis (All.) F.T. Hubb. [syn.]

Note: This is not a real synonym, but a late combination (T. Cope 8/2000, pers. comm.).

Eremobium lineare (Delile) Asch. & Schweinf. ex Boiss.; Fl.Pal.: Eremobium aegyptiacum (Spreng.) Boiss. var. lineare (Delile) Zohary [syn.]

Eremurus spectabilis M.Bieb. var. libanoticus (Boiss. & Blanche) O.Fedtsch.; Fl.Pal.: Eremurus libanoticus Boiss. & Blanche [syn.]

Erodium crassifolium L'Hér; Fl.Pal.: Erodium hirtum Willd. [syn.]

Erodium oxyrhynchum M.Bieb. subsp. bryoniifolium (Boiss.) Schoenb.-Tem.; Fl.Pal.: Erodium bryoniifolium Boiss. [syn.]

Erodium touchyanum Delile; Fl.Pal.: Erodium deserti (Eig) Eig [syn.]

Erophila praecox (Steven) DC.; Fl.Pal.: Erophila verna (L.) Besser [syn.]

Erucaria pinnata (Viv.) Täckh. & Boulos; Fl.Pal.: Erucaria uncata (Boiss.) Asch. & Schweinf. [syn.]

Erucaria rostrata (Boiss.) Greuter & Burdet; Fl.Pal.: Erucaria boveana Coss. [syn.]

Eryngium falcatum F.Delaroche; Fl.Pal.: Eryngium falcatum Laroche  [syn.]

Euphorbia grossheimii Prokh.; Fl.Pal.: Euphorbia isthmia Täckh. [syn.]

Euphorbia heterophylla L.; Fl.Pal.: Euphorbia geniculata Ortega  [syn.]